Proteins

2. Enzymes

2.6. Endopeptidase

Ep-A1d {894}. Isozyme 6. v: PI 294994 {894}.

Ep-D1e {894}. Isozyme 5. v: PI 294994 {894}.

2.20. Aromatic alcohol dehydrogenase

Aadh-A1.

Add at bottom of Aadh-A1 section: 'ma: XksuG44-5A (proximal) - 6.9 cM - Aadh-A1 - 24.7 cM - Xcdo412-5A (distal) {9959}.'

2.29. Starch branching enzyme
 
SbeI1 {9937}. 1DL {9937}. v: CS {9937}.
SbeI2 {9937}. 7BL {9937}. v: CS {9937}.

3. Endosperm Storage Proteins

3.1. Glutenins

At the end of the Glu-B1 section, delete

'Variation in the staining intensity of subunit 7 in different varieties has also been observed {1069}; possible low gene expression at Glu-B1 has been noted for Glu-B1w, where subunits 6*+8* stain very faintly {1146}.'

and substitute

'Variation in the staining intensity of subunit 7 in different varieties has also been observed {1069}; a duplication of the gene encoding subunit 7 has probably occurred in cultivar ‘Red River 68’, as evidenced by increased intensity of the subunit in SDS-PAGE and by approximately doubled intensity of restriction fragments carrying the gene in Southern blotting {9989}. Possible low gene expression at Glu-B1 has been noted for Glu-B1w, where subunits 6*+8* stain very faintly {1146}.'.

3.2. Gliadins
Revision of preamble:

Delete

'Variation at the Gli-1 loci was described earlier {634,996,1126} and applied in mapping experiments {1243,1125,196,422,1120}. A rational system of naming the alleles was produced by E.V. Metakovsky, N.I. Vavilov Institute of General Genetics, Moscow {988}. This nomenclature is reproduced below. Where two cultivars are given as prototypes for an allele, the first named is from the USSR and the second from elsewhere.'

and substitute

'Variation at the Gli-1 loci was described earlier {634,996,1126} and applied in mapping experiments {1243,1125,196,422,1120}. A rational system of naming the alleles has been produced by Dr. E.V. Metakovsky (Present Address: Unidad de Genética, Departamento de Biotecnologia, E.T.S.I. Agrónomos, Universidad Politecnica de Madrid, 28040 Madrid, Spain) {988}. This nomenclature is reproduced below. A considerable number of alleles have been added to the original list given in {988}, and referenced here accordingly. A few alleles have been deleted, because, following much detailed comparison, there is now doubt that they can be reliably distinguished from existing alleles {9981}; for the moment, however, the allelic letter in these cases has not been reused. To facilitate practical use of the list, the aim has been to give at least three standard cultivars from a range of countries for each allele {9981}. This has been achieved for the vast majority of entries and is a change from the original list compiled from {988}, where up to two standards were given. While the three or more standards described almost always include the original standards, some have been replaced for various reasons, such as international awareness of the cultivar, availability of seed, or the ease with which an allele can be identified in a particular genetic background {9981}. In the original list, where two cultivars were given as prototypes for an allele, the first named was from the USSR and the second from elsewhere; this is no longer the case, although pains have been taken to include a Russian cultivar where possible, to maintain the base of germplasm in which the alleles are available to be as wide as possible, as well as to acknowledge the research groups in the country where much of the pioneering work was carried out.

For discussion of null alleles at the Gli-1 and Gli-2 loci, see {9984}.'.

Delete the following from the preamble:

'A number of novel gliadin alleles were reported in {991}; they will be included in the next supplement to the catalogue.'

At the end of the preamble, delete

'The Gli-1 loci may be recognised by probes pcP387 {372} and pTag1436 {065}, and by specific microsatellite primers {252}.'

and substitute

'The Gli-1 loci may be recognised by probes pcP387 {372} and pTag1436 {065}, and by specific microsatellite primers {252}. Furthermore, it has been shown that probe pTag1436 differentiates gliadin alleles rather well; using this probe, families of gliadin alleles and some of their relationships have been described {9988}.'.

Gli-A1

Delete the previous corresponding entries and substitute the following:
 
Gli-A1a {988}. v: CS {988}; Castan {991}; Mentana {9986}; Mara {9986}.
Gli-A1b {988}. v: Bezostaya 1{988}; Mercia {988}; Tracy {991}.
Gli-A1c {988}. v: Ukrainka {988}; Gazul {9985}; Sava {994}.
Gli-A1d {988}. v: Dankowska {988}; Cabezorro {9985}.
Gli-A1e {988}. v: Open {991}; Touzelle {991}; Falchetto {988}.
Gli-A1f {988}. v: Maris Freeman {988}; Mironovskaya 808 {988}; Arminda {991}.
Gli-A1g {988}. v: Gabo {988}; Adalid {9985}.
Gli-A1h {988}. v: Sadovo I {988}; Predela {9981}; Krajinka {9981}.
Gli-A1i {988}. v: Saratovskaya 36 {988}.
Gli-A1j {988}. v: Lutescens 62 {988}.
Gli-A1k {988}. v: Courtot {991}; Soissons {991}; Spada {9986}; Skala (heterogeneous) {988}.
Gli-A1l {988}. v: Lesostepka 75 {988}; David {9986}; Salmone {9986}; Mura {9981}.
Gli-A1m {988}. v: Marquis {988}; Dneprovskaya 521 {988}; Carat {991}; Liocorno {9986}.
Gli-A1n {988}. v: Intensivnaya {988}.
Gli-A1o {988}. v: Cappelle-Desprez {991}; Capitole {991}; Oderzo {9986}; Odesskaya 16 (heterogeneous) {988}.
Gli-A1p {988}. v: Pyrotrix 28 {988}; Zagore {9981}.
Gli-A1q {988}. v: Akmolinka 1 {988}.
Gli-A1r {988}. v: Ranniaya 73 {988}; Barbilla {9985}.

Omission:

Gli-A1s.
Gli-A1s, not previously listed in the catalogue but reported in {9986}, is omitted from the catalogue temporarily because its status is in need of further confirmation {9981}.

Add:
 
Gli-A1t {9985}. v: Jeja del País {9985}.
Gli-A1u {9985}. v: Candeal Alcala {9985}.
Gli-A1null {9984,9987}. v: Saratovskaya 29 (mutant) {9987}; E.Mottin {9981}.

Gli-B1

Delete the previous corresponding entries and substitute the following:
 
Gli-B1a {988}. v: CS {988}.
Gli-B1b {988}. v: Bezostaya 1 {988}; Marquis {988}; Soissons {991}; Carat {991}; Liocorno {9986}.
Gli-B1c {988}. v: Siete Cerros 66 {988}; Prinqual {991}; Loreto {9986}.
Gli-B1d {988}. v: Chopin {991}; Dneprovskaya 521 {988}; Petrel {991}; Tiberio {9986}; Yécora {9985}.
Gli-B1e {988}. v: Lutescens 62 {988}; Apexal {991}; Fournil {991}; Oderzo {9986}.
Gli-B1f {988}. v: Maris Freeman {988}; Dankowska {988}; Mercia {988}; Cappelle-Desprez {991}; Capitole {991}.
Gli-B1g {988}. v: Galahad {988}; Sadovo 1 {988}; Champtal {991}; Tracy {991}; Mara {9986}.
Gli-B1h {988}. v: Krasnodonka {988}; Pepital {991}; Rudi {991}; Cabezorro {9985}.
Gli-B1i {988}. v: Insignia {988}; Ghurka {988}.
Gli-B1j {988}. v: Cluj 650 {988}.
Gli-B1k {988}. v: Mentana {9986}; Kremena {988}; De Carolis {9986}; Crvenkapa {994}.
Gli-B1l {988}. v: Clement {991}; Damier {991}; Fiocco {9986}.
Gli-B1m {988}. v: Pyrotrix 28 {988}; Et.d'Choisy {991}; Costantino {9986}.
Gli-B1n {988}. v: Intensivnaya {988}.
Gli-B1o {988}. v: Pippo {9986}; Levent {988}; Aragón 03 {9985}; San Rafael {9985}.
Gli-B1p {988}. v: New Pusa 834 {988}; Inia 66 {9985}.
Gli-B1q {9986}. v: Goelent {991}; Goya {991}; Gallo {9986}. 
Gli-B1r {995}. v: Gazul {9985}; Sevillano {9985}; Chinook {995}.
Add:
Gli-B1s {9986}. v: Salmone {9986}; Resistente {9986}; E.Mottin {9981}.
Omission:
Gli-B1t.
Gli-B1t, not previously listed in the catalogue but reported in {9986}, is omitted from the catalogue temporarily because its status is in need of further confirmation {9981}.
Add:
Gli-B1u {9985}. v: Negrillo {9985}.
Gli-B1v {9985}. v: Montjuich {9985}.
Gli-B1null{9984,9987,991}. v: Touzelle {991}; Florence Aurora {9985}.
Gli-D1
Delete the previous corresponding entries and substitute the following:
Gli-D1a {988}. v: CS {988}; Marquis {988}; Saratovskaya 36 {988}; Mentana {9986}; Prinqual {991}.
Gli-D1b {988}. v: Bezostaya 1 {988}; Galahad {988}; Cappelle-Desprez {991}; Et.d'Choisy {991}.
Gli-D1c {988}. v: Skorospelka Uluchshennaya (biotype) {988,9982}.
Gli-D1d {988}. v: De Carolis {9986}; Solo {988}.
Gli-D1e {988}. v: Gerek 79 {988}.
Gli-D1f {988}. v: Maris Freeman {988}; Gabo {988}; Carlos {991}; Orso {9986}.
Gli-D1g {988}. v: Ghurka {988}; Mironovskaya 808 {988}; Fournil {991}; Open {991}.
Gli-D1h {988}. v: Sadovo 1 {988}; Zlatostrui {9981}.
Gli-D1i {988}. v: Insignia {988}; Tselinogradka {988}; Napayo (biotype) {995}; San Rafael {9985}.
Gli-D1j {988}. v: Aubain {991}; Promin {988}; Petrel {991}; Inia 66 {9985}; Chinook {995}.
Gli-D1k {988}. v: Mara {9986}; Pippo {9986}; Kremena {988}; Cargimarec {991}.
Gli-D1l {988}. v: Longbow {988}; Artaban {991}; Corin {991}.
Add:
Gli-D1m {991}. v: Heurtebise {991}.
Gli-D1n {9981}. v: Blanquillo de Toledo {9985}.
Gli-D1null {9984,9987,991}. v: Darius {991}; Touzelle {991}; Saratovskaya 29 (mutant) {9987}.
Gli-A2
Delete the previous corresponding entries and substitute the following:
Gli-A2a {988}. v: CS {988}; Insignia {988}; Rieti DIV {9986}; Cabezorro {9985}.
Gli-A2b {988}. v: Bezostaya 1 {988}; Rivoli {991}; Tiberio {9986}; Aradi {9985}.
Gli-A2c {988}. v: Siete Cerros 66 {988}; Prinqual {991}; Loreto {9986}; Escualo {9985}.
Gli-A2d {988}. v: Dneprovskaya 521 {988}; Mocho Sobarriba {9985}.
Gli-A2e {988}. v: Sadovo 1 {988}; Cobra {991}; Mentana {9986}; Resistente {9986}; Sevillano {9985}.
Gli-A2f {988}. v: Maris Freeman {988}; Gala {991}; Sistar {9986}; Adalid {9985}.
Gli-A2g {988}. v: Cappelle-Desprez {991}; Ducat {988}; Mara {9986}; Mahissa 1 {9985}.
Gli-A2h {988}. v: Hereward {988}; Apollo {991}; N.Strampelli {9986}; Montjuich {9985}.
Gli-A2i {988}. v: Lesostepka 75 {988}.
Gli-A2j {988}. v: Recital {991}; Camp Remy {991}; Avalon {9981}; E.Mottin {9981}.
Gli-A2k {988}. v: Pyrotrix 28 {988}; Akmolinka 1 {988}; Estica {991}; Renan {991}; Zena {9986}.
Gli-A2l {988}. v: Longbow {988}; Champlein {991}; Chamorro {9985}.
Gli-A2m {988}. v: Marquis {988}; Rex {991}.
Gli-A2n {988}. v: Mironovskaya 808 {988}.
Gli-A2o {988}. v: Castan {991}; Touzelle {991}; Lontra {9986}; Calatrava {9985}.
Gli-A2p {988}. v: Capitole {991}; Pliska {988}; Clement {991}; S.Lorenzo {9986}; Cajeme 71 {9985}; Yecora {9985}.
Gli-A2q {988}. v: Saratovskaya 39 {988}; Montcada {9985}; Candeal Alcalá {9985}.
Gli-A2r {988}. v: Riband {988}; Open {991}; Genial {991}.
Gli-A2s {988}. v: Saratovskaya 36 {988}.
Gli-A2t {988}. v: Courtot {991}; Soissons {991}; Rinconada {9985}; Prostor {9981}.
Gli-A2u {988}. v: Titien {991}; Aragon 03 {9985}; Kirgizskaya Yubileinaya {988}; Saunders {995}.
Gli-A2v {988}. v: Kzul-Bas {988}.
Gli-A2w {988}. v: Bezenchukskaya 98 (biotype) {988}.
Gli-A2x {988}. v: Solo {988}.
Add:
Gli-A2y {9981}. v: Gentil Rosso 202 {9981}; PI 191245 {9981}.
Gli-A2z {9986}. v: Gallo {9986}; Giuliana {9986}.
Gli-A2aa {9985}. v: Navarro 122 {9985}.
Gli-A2ab {9985}. v: Navarro 150 {9985}.
Gli-A2null{9984,9987}. v: Saratovskaya 29 (mutant) {9987}.
Gli-B2
Delete the previous corresponding entries and substitute the following:
Gli-B2a {988}. v: CS {988}.
Gli-B2b {988}. v: Bezostaya 1 {988}; Cobra {991}; Sideral {991}; Gladio {9986}.
Gli-B2c {988}. v: Siete Cerros 66 {988}; Gabo {988}; Courtot {991}; Prinqual {991}; Loreto {9986}; Manital {9986}; Sinton {995}; Escualo {9985}; Yecora {9985}.
Gli-B2d {988}. v: Akmolinka 1 {988}; Tselinnaya 20 {988}; Friedland {991}.
Gli-B2e {9986}. v: Veronese {9986}; Arsenal {991}; Zlatna Dolina {994}.
Gli-B2f {988}. v: Maris Freeman {988}; Master {991}.
Gli-B2g {988}. v: Galahad {988}; Cappelle-Desprez {991}; Capitole {991}; Forlani {9986}.
Gli-B2h {988}. v: Mentana {9986}; Sadovo 1 {988}; Castan {991}; Sistar {9986}; Pane 247 {9985}; Partizanka {994}.
Gli-B2i {988}. v: Insignia {988}; Ghurka {988}.
Gli-B2j {988}. v: Farnese {9986}; Funo R250 {9986}; Novosadska Rana 1 {994}.
Gli-B2k {988}. v: Skala {988}.
Gli-B2l {988}. v: Longbow {988}; Tracy {991}; Clement {991}.
Gli-B2m {988}. v: Mironovskaya 808 {988}; Open {991}; Renan {991}.
Gli-B2n {988}. v: Solo {988}.
Gli-B2o {988}. v: Mara {9986}; Hardi {9981}; Rivoli {991}; Pippo {9986}; Slavianka {9981}; Odesskaya 16 {988}.
Gli-B2p {988}. v: Champtal {991}; Oderzo {9986}; Recital {991}; Gazul {9985}; Pliska {988}.
Gli-B2q {988}. v: Saratovskaya 39 {988}.
Gli-B2r {991}. v: Genial {991}; Arminda {991}; Jeja del País {9985}.
Gli-B2s {988}. v: Saratovskaya 36 {988}.
Gli-B2t {988}. v: Tselinogradka {988}.
Gli-B2u {988}. v: Kirgizskaya Yubileinaya {988}.
Gli-B2v {988}. v: Garant {991}; Libellula {9986}; Mahissa 1 {9985}; Poljarka {988}; Declic {991}.
Gli-B2w {995,9986}. v: Rieti DIV {9986}, Palata {9986}, Pembina {995}.
Add:
Gli-B2x {994}. v: Super Zlatna (biotype) {994}; Prostor {9981}; 251/83 {9981}.
Gli-B2y {9986}. v: Centauro {9986}; E.Morandi {9986}.
Gli-B2z {9985}. v: Maestro {9985}.
Gli-B2aa {9986}. v: Salmone {9986}; E.Mottin {9981}.
Omission:
Gli-B2ab.
Gli-B2ab, not previously listed in the catalogue but reported in {991}, is omitted from the catalogue because its status is in need of further confirmation {9981}.
Add:
Gli-B2ac {991}. v: Scipion {991}; Artaban {991}; Riol {991}; Lontra {9981}.
Gli-B2ad {991}. v: Champion {991}; Chopin {991}.
Gli-B2ae {991}. v: Priam {991}; Et.d'Choisy {991}; Campeador {9985}; Krajinka (biotype) {994}.
Gli-B2af {9985}. v: Montjuich {9985}; Mocho Sobarriba {9985}.
Gli-B2null {9984,9987}. v: Saratovskaya 29 {9987}.
Gli-D2
Delete the previous corresponding entries and substitute the following:
Gli-D2a {988}. v: CS {988}; Maris Freeman {988}; Tracy {991}; Sistar {9986}.
Gli-D2b {988}. v: Bezostaya 1 {988}; Cobra {991}; Farnese {9986}; Partizanka {994}.
Gli-D2c {988}. v: Siete Cerros 66 {988}; Eridano {9986}; Rieti DIV {9986}; Escualo {9985}.
Gli-D2d {988}. v: Dneprovskaya 521 {988}.
Gli-D2e {988}. v: Mironovskaya 808 {988}; Open {991}; Dollar {9985}; Lada {9981}.
Gli-D2f {988}. v: Rempart {991}; Créneau {991}; Kirgizskaya Yubileinaya {988}; Skorospelka Uluchshennaya {988}.
Gli-D2g {988}. v: Capelle-Desprez {991}; Futur {991}; Galahad {988}; Ghurka {988}; Mec {9986}.
Gli-D2h {988}. v: Capitole {991}; Garant {991}; Thatcher {995}; Chinook {995}; Sadovo 1 {988}.
Gli-D2i {988}. v: Insignia {988}; Lario {9986}.
Gli-D2j {988}. v: Mentana {9986}; Inia 66 {9985}; Gallo {9986}; Arcane {991}; Gazul {9985}.
Gli-D2k {988}. v: Skala {988}; Cabezorro {9985}; Crvencapa {994}.
Gli-D2l.
Gli-D2l is deleted from the catalogue because it has been shown not to demonstrate reliable differences compared with existing alleles {9981}.
Gli-D2m {988}. v: Marquis {988}; Rex {991}; Veronese {9986}; Yecora {9985}; Rinconada {9985}.
Gli-D2n {988}. v: Mercia {988}; Castan {991}; Pippo {9986}; Mahissa 1 {9985}; Champlein {991}.
Gli-D2o {988}. v: Omskaya 12 {988}.
Note: cultivars Salmone and Resistente, which carry Gli-D2aa {9981}, were erroneously given as standards for allele Gli-D2o in {9986}.
Gli-D2p {988}. v: New Pusa {988}.
Gli-D2q {988}. v: Soissons {991}; Fournil {991}; E.Mottin {9981}; Volshebnitsa (biotype) {988}.
Gli-D2r {988}. v: Mara {9986}; Montcada {9985}; Kremena {988}.
Gli-D2s {988}. v: Akmolinka1 {988}; Bezenchukskaya 98 {988}; Selkirk (biotype) {995}.
Add:
Gli-D2t {9986}. v: Golia {9986}; Gabo {9981}; Manital {9986}; Bokal {9981}.
Gli-D2u {9986}. v: Loreto {9986}; Martial {991}; Cibalka {9981}.
Gli-D2v {991}. v: Epiroux {991}; Arbon {991}.
Gli-D2w {9985}. v: Navarro 150 {9985}; Javelin {9981}; Hopps {9981}; Canaleja {9985}.
Gli-D2x {9985}. v: Montjuich {9985}; Blanquillo {9985}.
Gli-D2y {9985}. v: Candeal Alcalá {9985}.
Gli-D2z {9985}. v: Aragon 03 {9985}.
Gli-D2aa {9981}. v: Salmone {9981}; Resistente {9981}.
Gli-D2null {9984,9987}. v: Saratovskaya 29 (mutant) {9987}.
Gli-A3
Add:
Gli-A3a {9983}. v: CS, Prinqual, Courtot, Tselinogradka, Bezenchukskaya 98.
Gli-A3b {9983}. v: Bezostaya 1.
Gli-A3c {9983}. v: Anda.
Gli-A3d {9983}. v: Saratovskaya 210, Kharkovskaya 6, Richelle.

Each of the above Gli-A3 alleles, apart from Gli-A3d, which is a null, controls one minor omega-gliadin with molecular mass about 41k that occurs in the middle of the omega-region of APAGE fractionation. Gliadins controlled by these alleles differ in their electrophoretic mobility in APAGE in that the fastest of three known Gli-A3-gliadins is controlled by Gli-A3a and the slowest by Gli-A3c {9983}.

In the paragraph that begins,

'It is not clear how Gli-Sl4 and Gli-Sl5 relate to ...',

delete

'It has yet to be established whether this is one of a series of orthologous loci.
 
  Gli-A4 {1205}. 1AS {1205}. v: Perzivan biotype 2.

Dubcovsky et al. {277} did not find evidence for the simultaneous presence of both Gli-A3 and Gli-A4 in five 1A or 1Am mapping populations and concluded that Gli-A4 should be considered to be Gli-A3 until conclusive evidence for the former is obtained.'

and substitute

'However, Metakovsky et al. {9983} have since shown that this locus and Gli-A3 are, in fact, the same locus. Furthermore, Dubcovsky et al. {277} did not find evidence for the simultaneous presence of both Gli-A3 and Gli-A4 in five 1A or 1Am mapping populations and concluded that Gli-A4 should be considered to be Gli-A3 until conclusive evidence for the former is obtained. For these reasons, the locus Gli-A4 is deleted from the catalogue.'.

In the paragraph that begins,

'A locus designated Gli-5 controlling omega-gliadins was mapped ....',

delete

'An estimate for the map distance between Gli-A5 and Gli-A1 was not reported, although evidence was provided that the linkage is of a similar order of magnitude to the Gli-B5 - Gli-B1 distance. Although no orthologous locus was reported for chromosome 1D, the authors cited studies {992,987} reporting a recombination distance of 1 % between two gliadin loci on chromosome 1D, which they considered may have been due to the presence of a locus on 1D orthologous to Gli-B5.'

and substitute

'An estimate for the map distance between Gli-A5 and Gli-A1 was not reported, although evidence was provided that the linkage is of a similar order of magnitude to the Gli-B5 - Gli-B1 distance, and, since then, Gli-A5 has been shown to be tightly linked (0-2%) to Gli-A1 {9983}. Although no orthologous locus was reported for chromosome 1D, the authors cited studies {992,987} reporting a recombination distance of 1 % between two gliadin loci on chromosome 1D, which they considered may have been due to the presence of a locus on 1D orthologous to Gli-B5.'.

Gli-A5

Add:
 
  Gli-A5a {9983}.   v: CS.
  Gli-A5b {9983}.   v: Marquis.

Gli-A5a is null. Allele Gli-A5b controls two slow-moving, easily-recognizable omega-gliadins. It is present in all common wheat cultivars having alleles Gli-A1m and Gli-A1r (and, probably, in those having Gli-A1e, Gli-A1l and Gli-A1q), because earlier (for example, in {988}) two minor omega-gliadina encoded by Gli-A5b were considered to be controlled by these Gli-A1 alleles {9983}.

Gli-B5

Add:
 
  Gli-B5a {1147}.   v: CS.
  Gli-B5b {1147}.   v: Salmone.

The situation is fully analogous with Gli-A5: There are currently two alleles differing in the presence/absence of two minor omega-gliadins. Gli-B5a is null. In {988}, omega-gliadins controlled by Gli-B5 (allele Gli-B5b) were attributed to alleles at the Gli-B1 locus (alleles Gli-B1c, i, k, m, n and o).

Add:
 
Gli-A6 {9983,993}. 1AS {9983}.    
  Gli-A6a {9983}.   v: CS, Bezostaya 1.
  Gli-A6b {9983}.   v: Bezenchukskaya 98.
  Gli-A6c {9983}.   v: Courtot, Anda, Mironovskaya 808.

Gli-A6 was first explicitly described in {9983}, but it is now known that it was first observed without designation in {993}. There is strong evidence that it is distinct from Gli-A3 and Gli-A5, mapping distally to Gli-A1, with which it recombines at a frequency of 2-5%. Currently three alleles are known, of which Gli-A6c is particularly well-described in {9983}: the molecular mass of the omega-gliadin controlled by this allele is slightly lower than those of the omega-gliadins controlled by Gli-A3 alleles. In {988}, the omega-gliadin controlled by Gli-A6c was attributed to Gli-A1f. Gli-A6c is rather frequent in common wheat and may relate to dough quality (preliminary data {9983}). Gli-A6a is null {9983}.

Delete from end of gliadin section:

'The Gli-1 loci may be recognised by pcP387 {372}, pTag1436 {066} and by specific microsaellite primers {252}.'

5. Other proteins
5.1. Lipopurothionins

Pur-A1.

Add at end of section: 'A PCR marker specific for Pur-A1 was developed in {9976}.'

Pur-B1.

Add at end of section: 'A PCR marker specific for Pur-B1 was developed in {9976}.'

Pur-D1.

Add at end of section: 'A PCR marker specific for Pur-D1 was developed in {9976}.'

Pur-R1.

Add at end of section: 'A PCR marker specific for Pur-R1 was developed in {9976}.'

5.6. Waxy Proteins

Add: 'Lists of cultivars, lines and landraces of tetraploid and hexaploid wheats with different, mostly null, alleles at the Wx loci are given in {9910,9911,9912,1053,1054,9913,9915,9916,1650,9917}.'