2.10. Hexokinase
 
Hk-B1 {006}.   1BS {006}. v: CS.
Hk-D1 {006}.   1DS {006}. v: CS.
Hk-A2 {810}.   3A {810}. v: CS.
Hk-A2a {810}.
    v: CS.
Hk-A2b {810}.
s: CS*/{Sears' Synthetic 3A.
v: Sears' Synthetic IPSR1190903.
Hk-B2 {006}.   3BS {006,810}. v: CS.
Hk-D2 {810}.   3DS {810}. v: CS.
Hk-D2a {810}.
    v: CS.
Hk-D2b {810}.
    v: Sears' Synthetic IPSR1190903.
Hk-E2 {006}.   3ES {006}. ad: CS/E. elongata.

Allelic variation was observed in three of 55 hexaploid accessions {006}.

2.11. Lipoxygenase
 
Lpx-A1 [{516}]. [Lpx-B1 {516}]. 4AL {516}. v: CS.
Lpx-B1 [{516}]. [Lpx-A1 {516}]. 4BS {516}. v: CS.
Lpx-B1a [{1533}].
[Lpx-A1a {936}].   v: CS.
Lpx-B1b [{1533}].
[Lpx-A1b {936}].   v: Bosanka {1533}.
Lpx-D1 {516}.   4DS {516}. v: CS.
Lpx-E1 {518}.   4ES {518}. ad: CS/E. elongata.
Lpx-H1 {716}.   4H {716}. ad: CS/Betzes.
Lpx-A2 {516}.   5AL {516}. v: CS.
Lpx-B2 {516}.   5BL {516}. v: CS.
Lpx-D2 {516}.   5DL {516}. v: CS.
Lpx-E2 {518}.   5EL {518}. ad: CS/E. elongata.
Lpx-V2 {242}.   5V. ad: CS/D. villosum.
Lpx-H2 {716}.   5H {716}. ad: CS/Betzes.
Lpx-Ss2 {1140}.   5Ss {1140}. ad: CS/T. searsii.

2.12. Malate dehydrogenase
 
Mdh-A1. [Mdh2A {087}]. 1AL {087}. v: CS.
Mdh-B1. [Mdh2B {087}]. 1BL {101,087}. v: CS.
Mdh-D1. [Mdh2D {087}]. 1DL {087}. v: CS.
Mdh-H1 {1153}.   1HL {1153}. ad: CS/Betzes.
Mdh-Hch1 {352}.   1Hch {352}. ad: MA/H. chilense.
Mdh-R1. [Mdh2-1 {1252}]. 1RL {1252}. ad: CS/Imperial 1R; Kharkov/Dakold 1R; Holdfast/King II 1RL.
Mdh-Ss1 {1140}.    1Ss {1140}. ad: CS/T. searsii.
Mdh-H2. [Mdh2-b2 {090}]. 3H {090}.  
Mdh-R2. [Mdh2-2 {1252}]. 3R {1252}. ad: CS/Imperial.

A third set of dimeric MDH isozymes identified in mature grain are separable from MDH-1 and MDH-2 by their higher pI's in IEF {811}.
 
Mdh-A3 {811}.    5AS. v: CS.
Mdh-A3a {811}.
    v: CS.
Mdh-A3b {811}.
    v: Bersee.
Mdh-B3 {811}.    5BS. v: CS.
Mdh-B3a {811}. 
    v: CS.
Mdh-B3b {811}.
    v: Hope.
Mdh-D3 {811}.    5DS. v: CS.
Mdh-D3a {811}.
    v: CS.
Mdh-D3b {811}.
    v: Sears' Synthetic.
Mdh-E3 {811}.   5ES. ad: CS/Ag. elongatum.
Mdh-H3 {811}.   5H. ad: CS/Betzes.
Mdh-U3 {811}.   5U. ad: CS/Ag. umbellulatum.
Mdh-R4 {360}.   1RL {360}. v: Various crosses.

2.13. Peroxidase

Peroxidase (EC1.11.1.7) isozymes have high tissue specificity. Staining and electrophoretic systems are reviewed in 90. PER-1, -2, -3, -4 and -5 are all reported in {816}.

PER-1 is expressed in leaf {012} and coleoptile {816} tissue.
 
Per-B1 {012}.   1BS {012,919}. v: CS.
Per-D1 {012}.   1DS {012,919}. v: CS.
Per-D1a {012}.
    v: CS.
Per-D1b {012}.
    v: Sears' Synthetic.
Per-Hch1 {012}.   1Hch {012}. ad: CS/H. chilense.
Per-R1 {012}. [Prx {1561}]. 1RS {012,1561}. ad: CS/King II {012}; Holdfast/King II {1561}.
tr: Veery 'S' {012}.
Per-V1 {241}.   1V {241}. ad: Creso/D. villosum.

PER-2 is expressed in young leaf {118}, coleoptile and root {816} tissue.
 
Per-A2.   2AS. v: CS.
Per-A2a {816}.
    v: CS.
Per-A2b {816}.
    v: Timstein.
Per-B2 {118}.   2BS {118}. v: CS.
Per-B2a {816}.
    v: CS.
Per-B2b {816}.
    v: Sears' Synthetic IPSR1190903.
Per-D2 {118}.   2DS {118}. v: CS.
Per-H2 {118}. [Per-5 {095}]. 2H {118}. ad: CS/Betzes.
Per-R2 {118}.   2RS {118}. ad: CS/Imperial; Kharkov/Dakold.

PER-3 is expressed in embryo {119,816} and scuteller {119} tissue.
 
Per-A3 {119}.   3AL {119}. v: CS.
Per-A3a {816}.
    v: CS.
Per-A3b {816}.
    v: Timstein.
Per-A3c {816}.
    v: Hobbit 'S'.
Per-B3 [{086}], {119}. [Per4 {961}]. 3BL{086,119}. v: CS.
Per-B3a {816}.
    v: CS.
Per-B3b {816}.
    v: Hope.
Per-B3c {816}.
    v: T. macha IPSR1240005.
Per-B3d {816}.
    v: Timstein.
Per-B3e {816}.
    v: Sears' Synthetic IPSR1190903.
Per-D3 [{086}],{119}. [Per5 {961}]. 3DL{086,119}. v: CS.
Per-D3a {816}.
    v: CS.
Per-D3b {816}.
    v: Hope.
Per-D3c {816}.
    v: Timstein.
Per-D3d {816}.
    v: T. macha IPSR142005.
Per-D3e {816}.
    v: Sava.
Per-D3f {816}.
    v: Cheyenne.
Per-D3g {816}. 
v: Sears' Synthetic
IPSR1190903.

Varietal variation for Per-3 was reported in {094}.

PER-4 is expressed in endosperm tissue {086,119}.
 
Per-A4 [{695}], {119}. [Per3 {961}]. 7A {695}, 7AS {694,086,119}. v: CS.
Per-A4a {816}.
    v: CS.
Per-A4b {816}.
    v: Hope.
Per-A4c {816}.
    v: Sicco.
Per-B4 [{695}], {119}. [Per2 {961}]. 4A {695}, 4AL {086,119,694}. v: CS.
Per-B4a {816}.
    v: CS.
Per-B4b {816}.
    v: Hope.
Per-B4c {816}.
    v: Thatcher.
Per-D4 [{695}], {119}. [Per1 {961}]. 7D {695}, 7DS {694,086,119}. v: CS.
Per-D4a {816}.
    v: CS.
Per-D4b {816}.
    v: Thatcher.
Per-Age4.   7AgeS {694}. tr: Certain CS 7D/Age lines.
Per-Agi4. [Per-Agi3 {374}]. 7Agi {168}. ad: Vilmorin 27/Ag. intermedium.

Cultivar variation for Per-4 was reported in {094}.

PER-5 is expressed in roots {816}.
 
Per-D5 {816}.   2DS {816}. v: CS.
Per-Sl5 {816}.
  2Sl {816}. ad: CS/Ae. longissima.

2.14. Phosphodiesterase
 
Pde-A1 [Pde-A3 {1590}]. 3A {1590}, 3AS {1589}. v: CS.
Pde-B1. [Pde-B3 {1590}]. 3B {1590}, 3BS {1589}. v: CS.
Pde-D1. [Pde-D3 {1590}]. 3DS {1590}. v: CS.
Pde-Sl1.   3SlS {172}. ad: CS/T. longissimum.
Pde-V1 {1518}.   3VS {1518}. ad: CS/D. villosum.

2.15. Phosphogluconate dehydrogenase
 
Pgd1 [{282}]. [Pgd3 {282}, Pgd-A3 {963}]. 7AmS {282}. v: T: monoccum {664}
PgdR1.   4RL {1191}. ad: CS/Imperial; Holdfast/King II.
PgdR2.   6RL {1191}. ad: CS/Imperial; Holdfast/King II.

Loci were also identified in 6B {1435}, 1EL {1435}, 1HL {147,1072}, 1Hch {352} and 1RL {779}.

2.16. Phosphoglucomutase
 
Pgm-A1 {088}. [Pgm-B1 {088}]. 4AL {088}. v: CS.
Pgm-D1 {088}.   4DS {088}. v: CS.
Pgm-Agi1 [{361}], {374}. [Pgm-X1 {361}]. 4Agi {361}. ad: Vilmorin 27/Ag. intermedium.
Pgm-H1. [Pgm-b1 {090}]. 4H {090}. ad: CS/ Betzes.
Pgm-Hch1 {351}.   4Hch {351}. ad: MA/H. chilense.
Pgm-R1 4RS {1561,1253}. ad: CS/Imperial 4RS
{1253,1561};
Kharkov/Dakold 4R
{1253}; Holdfast/King II
4RS {1253,1561}.

2.17. Shikimate dehydrogenase
 
Skdh-A1 {706,1065}.   5AS {706,1065}. v: CS.
Skdh-B1 {706,1065}.   5BS {706,1065}. v: CS.
Skdh-D1 {706,1065}.   5DS {706,1065}. v: CS.
Skdh-H1 {085}.   5H {085}. ad: CS/Betzes.
Skdh-Mv1 [{985}].  [Skdh-Mv1 {985}]. 5Mv. su: 5Mv(5A), 5Mv(5D).
Skdh-R1 {706}. 5RS {706}, 5R {085}. ad: CS/King II {085}; CS/Imperial {706}; Kharkov/Dakold {085}.
tr: CS 4AS-5RL; CS 5BL-5RL.
Skdh-Sl1 {085}.   5SlS {085}. ad: CS/ Ae. longissima.
Skdh-Ss1 {1140}.   5Ss {1140}. ad: CS/T. searsii.
Skdh-V1 {085}.   5V {085}. ad: CS/D. villosum.
Skdh-U1.   5U {706}. ad,su: CS/Ae. umbellulata.
Skdh-Ht1 {1037}.   5Ht {1037}.  ad: CS/E. trachycaulus.

2.18. Superoxide dismutase
 
Sod-A1 {1066}.   2AL {1066}. v: CS.
Sod-B1 {1066}.   2BL {1066}. v: CS.
Sod-D1 {1066}.   2DL {1066}. v: CS.
Sod-E1 {808}.   VI E {808}. ad: CS/Ag. elongatum.
Sod-H1 {716}.   2H {716}. ad: CS/Betzes.
Sod-R1 {1066}. [Sod-3 {586}]. 2R {1066}. ad: CS/Imperial.
Sod-Ss1 {1140}.   2Ss {1140}. ad: CS/T. searsii.
Sod-VI {1026}.   7V {1026}. ad: CS/D. villosum.

2.19. Triosephosphate isomerase
 
Tpi-A1 {1139}.   3AS {1139}. v: CS.
Tpi-B1 {1139}.   3BS {1139} v: CS.
Tpi-D1 {1139}.   3DS {1139}. v: CS.
Tpi-E1 {1139}.   3E {1139}. ad: CS/E. elongata.
Tpi-H1 {1139}.   3H {1139}. ad: CS/Betzes.
Tpi-R1 {1139}.   3R {1139}. ad: CS/Imperial; Kharkov/Dakold.
Tpi-Sl1 {1139}.   3Sl {1139}. ad: CS/ T. longissimum.
Tpi-A2 {1139}.   5AL {1139}. v: CS.
Tpi-B2 {1139}.   5BL {1139}. v: CS.
Tpi-D2 {1139}.   5DL {1139}. v: CS.
Tpi-H2 {1139}.   5H {1139}. ad: CS/Betzes.
Tpi-R2 {1139}.   5R {1139}. ad: CS/Imperial; Kharkov/Dakold.
Tpi-Sl2 {1139}.   5Sl {1139}. ad: CS/Ae longissima.
Tpi-U2 {1139}.   5U {1139}. ad: CS/Ae umbellulata.
Tpi-Agi2 {374}. 5Agi {374}. ad: Vilmorin 27/Ag.
intermedium.

2.20. Aromatic alcohol dehydrogenase
 
Aadh-A1. [Adh-A2 {584}]. 5AL {584}. v: CS.
Aadh-A1a.     v: CS; 133 other accessions {584}.
Aadh-A1b.     v: T. spelta; K-24696; other accessions {584}.
Aadh-B1. [Adh-B2 {584}]. 5BL {584}. v: CS.
Aadh-B1a.     v: CS {1533}.
Aadh-B1b.     v: Drina {1533}.
Aadh-C1 [{1278}].   C {1278}. ad: T. aestivum cv. Alcedo/Ae. caudata line C.
Aadh-D1. [Adh-D2 {584}]. 5DL {584}. v: CS.
Aadh-E1. [Adh-E2 {518}]. 5EL {518}. ad: CS/E. elongata.
Aadh-R1.   5RL {1280}. ad: Holdfast/King II.
Aadh-A2. [Adh-A3 {508}]. 6A {1279}, 6AL {513,587}. v: CS {513}; Carola {1279}.
Aadh-B2. [Adh-B3 {508}]. 6B {1279}, 6BL {513}. v: CS {513}; Carola {1279}.
Aadh-D2. [Adh-D3 {508}]. 6D {1279}, 6DL {513}. v: CS {513}; Carola {1279}.
Aadh-Age2 {1575}.   6Age {1575}. ad,su: Rescue/Ag. elongatum.
Aadh-E2. [Adh-E3 {518}]. 6Eß {518}. ad: CS/E. elongata.
Aadh-R2.   6RL {1280}. ad: Holdfast/King II.
Aadh-V2 {241}.   6V {241}. ad: CS/D. villosum.

The Aadh-1 and Aadh-2 loci were designated with the synonyms Adh-2 and Adh-3, respectively, in a

number of publications in addition to {508,518,584}. These include: {510,509,511,519,517,587,1066,1139}.

2.21. Aconitase
 
Aco-A1 {189}.   6AL {189}. v: CS.
Aco-A1a.
    v: CS {1533}.
Aco-A1b.
    v: Dubravka {1533}.
Aco-B1 {189}.   6BL {189}. v: CS.
Aco-B1a.
    v: CS {1533}.
Aco-B1b.
    v: Dubravka {1533}.
Aco-B1c.
    v: Slavonka {1533}.
Aco-D1 {189}.   6DL {189}. v: CS.

Further alleles at Aco-A1 and Aco-B1 are listed in {1127}; these have not been tested against those found in {1533}.
 
Aco-Age1 {1575}.   6Age {1575}. ad,su: Rescue/Ag. elongatum.
Aco-E1 {189}.   6Eß {189}. ad: CS/E. elongata.
Aco-H1 [{147}],{189}. [Aco-1 {147}]. 6H {147}, 6HL {189}.  ad: CS/Betzes.
Aco-R1 {189}.   6RL {189}. ad: Sturdy/PI 252003.
Aco-Sl1 {189}.   6Sl {189}. ad: CS/Ae. longissima.
Aco-Ss1 {1140}.   6Ss {1140}. ad: CS/T. searsii.
Aco-U1 {189}.   CSU-31 {189}. ad: CS/Ae. umbellulata.
Aco-A2 {189}.    5AL{189}. v: CS.
Aco-B2 {189}.   4BL {1513}. v: CS.
Aco-B2a {1513}.
    v: CS.
Aco-B2b {1513}.
    v: PI 278437.
Aco-B2c {1513}.
    v: PI 182575.
Aco-B2d {1513}.
    v: PI 157589.
Aco-D2 {189}.   4DL {1513}. v: CS.
Aco-E2 {189}.   4EL {189}. ad: CS/E. elongata.
Aco-Mv2 [{1341}]. [Aco-Mv2 {985}]. 5Mv. su: 5Mv(5A),5Mv(5D).
Aco-R2 {189}.   5RL {189}. ad: CS/King II 5R; Holdfast/ King II 5RL.
Aco-Ss2 {1140}.   4Ss {1140}. ad: CS/T. searsii.

2.22. NADH dehydrogenase
 
Ndh-A1 [{513}], {1037}. [Ndh-B1 {513}]. 4AL {513}. v: CS.
Ndh-A1a [{1533}].
[Ndh-B1a {936}].   v: CS.
Ndh-A1b [{1533}].
[Ndh-B1b {936}].   v: Sutjeska.
Ndh-A1c [{11240].
[Ndh-B1c {936}].   v: Fruskogorka.
Ndh-A1d [{1037}].
[Ndh-A1b {1037}].   v: Hope, Timgalen.
Ndh-B1 [{513}]. [Ndh-A1 {513}]. 4BS {513}. v: CS.
Ndh-D1 {513}.   4DS {513}. v: CS.
Ndh-E1 {362}.   4E {362}. ad: CS/Ag. elongatum.
Ndh-H1 [{147}], {513}. [Nadhd-1 {147}]. 4H {147}, 4HS {813}. ad: CS/Betzes.
Ndh-V1 {241}.   4V {241}. ad: CS/D. villosum.
Ndh-Hch1 {813}.   4Hch {813}. ad: CS/H. chilense.
Ndh-R1 {813}.   4R {362}, 4RS {813}. ad: CS/Imperial, CS/King II {813,362}. CS/Dakold {362}.
Ndh-S11 {813}.   4Sl {813}. ad: CS/Ae. longissima.
Ndh-U1 {362}.   A {362}. ad: CS/Ae. umbellulata.
Ndh-A2 {813}.   7A {813}. v: Hope.
Ndh-D2 {813}.   7DS {813}. v: CS.
Ndh-R2 {813}.   7RS {813}. ad: CS/Imperial, CS/King II, Holdfast/King II (7R).
Ndh-A3 {813}.   3AL {813}. v: CS.
Ndh-B3 {813}.   3BL {813}. v: CS.
Ndh-B3a {813}.
    v: CS.
Ndh-B3b {813}.
    v: Carmen.
Ndh-D3 {813}.   3DL {813}. v: CS.
Ndh-H3 {813}.   3HL {813}. ad: CS/Betzes.
Ndh-R3 {813}.   6RL {813}. ad: Holdfast/King II, CS/Imperial (6R), CS/King II (6R).
Ndh-Sl3 {813}. 3SlL {813}. ad: CS/Ae. longissima.
ad: CS/Ae. sharonesis (3Sl).
Ndh-A4 {813}.   3AS {813}. v: CS.
Ndh-B4 {813}.   3BS {813}. v: CS.
Ndh-H4 {813}.   3HS {813}. ad: CS/Betzes, CS/King II.
Ndh-E4 {813}.   3ES {813}. ad: CS/Ag. elongatum.
Ndh-R4 {813}.   3RS {813}. ad: CS/King II, CS/Imperial (3R).

2.23. Dipeptidase
 
Dip-A1 [{454}], {700}.  [Pept-A1 {454}]. 6AL {454,700}. v: CS.
Dip-A1a {700}
    v: CS.
Dip-A1b {700}.
    v: Cheyenne.
Dip-B1 [{454}], {700}. [Pept-B1 {1533}]. 6BL {454,700}. v: CS.
Dip-B1a {700}.
    v: CS.
Dip-B1b {700}.
    v: Cappelle-Desprez.
Dip-D1 {700}.   6DL {700}. v: CS.
Dip-H1 [{147}], {700}.  [Pept-1 {147}, Dip 1 {145}]. 6H {147,145,700}. ad: CS/Betzes.
Dip-J1 {700}.   6J {700}. ad: CS/Ag. junceum.
Dip-V1 {700}.   6V {700}. ad: CS/D. villosum.

2.24. Malic enzyme

A dimeric enzyme extractable from mature grains.
 
Mal-A1 {809}.   3AL. v: CS.
Mal-B1 {809}.   3BL. v: CS.
Mal-B1a {809}.
    v: CS.
Mal-B1b {809}.
    v: T. spelta IPSR line 1.
Mal-B1c {809}.
    v: Sears' Synthetic.
Mal-D1 {809}.   3DL. v: CS.
Mal-E1 {809}.   3E. ad: CS/Ag. elongatum.
Mal-H1 {809}.   3H. ad: CS/Betzes.
Mal-R1 {809}.   3R. ad: CS/Imperial.

2.25. Adenylate kinase
 
Adk-A1 {091}.  [Adk-a {091}]. 7AL {091}. v: CS.
Adk-B1 {091}. [Adk-b {091}]. 7BL {091}. v: CS.
Adk-D1 {091}.  [Adk-d {091}]. 7DL {091}. v: CS.
Adk-E1 {091}.   7E {091}, 7E {1435}. ad: CS/E. elongata.
Adk-H1 {091}.   7H {091}, 7HS {1435}. ad: CS/Betzes.
Adk-Mv1 [{985}]. Adk-Mv1 {985}]. 7MvL. su: 7Mv{7D}.
Adk-R1 {091}.   7RL {091}. ad: CS/Imperial; Holdfast/King II.
Adk-U1 {091}.   E {091}. ad: CS/Ae. umbellulata.
Adk-AgiI {091}.   7Agi {091}. ad: CS/Ag. intermedium.
Adk-H2.   6HL {1435}. ad: CS/Betzes.

2.26. Glutamate-pyruvate transaminase
 
Gpt-A1 {1435}.   1AS {1435}. v: CS.
Gpt-B1 {1435}.   1BS {1435}. v: CS.
Gpt-D1 {1435}.   1DS {1435}. v: CS.
Gpt-E1 {1435}.   1ES {1435}. ad: CS/E. elongata 1E.
Gpt-H1 {1435}.   1H {1435}. dv: H. vulgare cv. Betzes.

2.27. Catalase
 
Cat-B1 {1466}.  [Cat-A1 {1466}]. 4BL {1466}. v: CS.

2.28. ß-glucosidase
 
ß-Gls {282}.   2AmL {282}. dv: DV92.
ß-Glsa {282}.
    dv: DV92.
ß-Glsb {282}. 
  Null allele. dv: G3116.