1994 MISSOURI WINTER WHEAT PERFORMANCE TESTS K. D. Kephart, A. L. McKendry, D. N. Tague and J. E. Berg Introduction Genetic improvement of wheat varieties has contributed about 40 to 50 percent of the total improvement in wheat yields attained over the past 50 years. Both public and private wheat breeding programs are constantly striving towards greater yield potential, improved quality and better host plant resistance to disease and insect pests in the new varieties released for commercial production. Over 100 new soft red winter wheat varieties alone have been released to U.S. farmers in the last 10 years. Wheat growers are reluctant to adopt new varieties without adequate information concerning adaptation and performance. The objective of the Missouri Winter Wheat Performance Tests is to provide wheat growers in Missouri with a reliable, unbiased, up-to- date source of information that will permit valid comparisons among improved wheat varieties. This information should help Missouri wheat growers select varieties best suited to their particular area and growing conditions. This report summarizes soft and hard red winter wheat variety trials conducted throughout Missouri during the 1993-94 cropping season. Variety Testing Procedures Locations --------- The soft red winter wheats were planted at seven locations throughout the state (Fig. 1) including Portageville and Charleston in the southeast, Mt. Vernon and Lamar in the southwest and Columbia, Novelty and in Grundy County near Trenton in the northern region of the state. The hard red winter wheats were planted at three sites including Columbia, Mt. Vernon, and in Grundy County near Trenton. ---------------------------------------------------------------------- +--------------------+ \ Trenton Novelty > * * \ \ \ \ \ | Columbia V\ | * | | < | \ | \ | * Lamar \ | * Mt. Vernon *< Charleston | > +----------------------, / / */ Portageville ----- Figure 1. Missouri winter wheat test locations in 1993. ---------------------------------------------------------------------- Entries and Seed Sources ------------------------ In 1994, 63 soft red and 5 hard red winter wheats were tested in Missouri. The soft red winter wheats were comprised of 13 public varieties, 8 public experimental entries and 42 proprietary varieties. The hard red winter wheats were comprised of 3 public varieties and 2 proprietary entries. Public varieties adapted to Missouri growing conditions or recommended by the state of origin were entered into the appropriate variety test. Numbered entries preceded by a state designation (e.g. MO12258, KY83C-16-2) are experimental lines provided by the foundation seed organization or wheat breeder of the originating state. Named public varieties were acquired from the foundation seed organization of the originating state or from the University of Missouri Foundation Seed Program. Names of commercially available entries evaluated in 1994 and their seed sources are listed in Table 1. Proprietary entries are submitted for testing on a fee basis by their owners or sponsors. Condition of the seed lot used in these tests for each entry is the responsibility of the company or organization sponsoring that entry. Experimental Design and Seeding Methods --------------------------------------- Each soft red winter wheat experiment was planted using a 8x8 lattice design with four replications. Hard red winter wheats were arranged in a randomized complete block design replicated 4 times. Test plots consisted of a 15 foot, 6-row plot with 7-inch row spacing. All entries were seeded at approximately 1.5 million seeds per acre, roughly equivalent to seeding 1.5 to 2 bushels per acre. Actual seeding rates were calculated from the thousand kernel weights determined for each entry (Table 2) and varied from 60 to 151 pounds per acre. Seeding rates were not adjusted for germination. Except for the Trenton location, all entries were seeded 1.25 to 1.5 inches deep into conventional seedbeds using a plot drill equipped with conventional double-disk openers. At the Trenton site, all entries were no-till seeded directly into soybean stubble using a plot drill equipped with AcraplantTm no-till openers. Agronomic Practices ------------------- Basic agronomic practices are given in Table 3 by location. Planting dates were based on Hessian fly-free dates recommended for each location. Nitrogen was applied in split fall/spring applications. Spring nitrogen applications were generally made after initial green up. Preplant phosphorous and potassium applications were based on soil test recommendations provided by the University of Missouri Soil Testing Laboratory located at Columbia. Description of Data Collected Yield ----- All rows of each test plot were trimmed 26 inches and harvested using an experimental plot combine. Recorded grain yields were adjusted to 13% grain moisture content, and are reported in bushels per acre based on a 60 pound per bushel test weight. In addition to yields obtained in 1994, two (1993-94) and three (1992-94) year averages are provided for both soft red and hard red winter wheat entries tested during previous cropping seasons. Test Weight and Grain Moisture Content -------------------------------------- Test weight (pounds per bushel) and percent grain moisture content were obtained for each plot using a Dickey-john GAC II grain analyzer. Plant Height ------------ Plant height was measured in inches from the soil surface to the top of the head, excluding the awns if present. Reported values have been rounded to the nearest inch. Lodging ------- Lodging severity was rated at locations where lodging was significant. Plots were rated on a severity scale of 0 to 9 where 0 = no lodging and 9 = plants completely flat. Winter Survival --------------- Percent winter survival was estimated for each plot after initial spring green-up at locations where significant winter injury occurred. In 1994, differences in winter survival among varieties were noted at all locations. Heading Date ------------ Heading was noted when 50% of the heads in a plot had extended above the flag leaf collar. Heading dates were recorded in Julian days (number of days from January 1) for statistical purposes. Corresponding calendar dates also are presented. Disease Ratings --------------- In 1994, all soft red winter wheat entries were tested for seedling reactions to Septoria leaf blotch under greenhouse conditions. Four replications of ten plants per entry were inoculated with S. tritici spores at the 3-leaf stage of development. After four weeks, a necrosis rating (0=no injury, 9=total death) was determined for each entry (Table 5). Resistance ratings are not given, but higher necrosis ratings are indicative of greater susceptibility to Septoria leaf blotch during the seedling stages of development. Some results may be inconsistent with known field reactions to Septoria leaf blotch. In some cases, these results may have been biased by use of triadimenol (Baytan) fungicide as a seed treatment on the seed submitted for testing. The occurence of foliar diseases was light to modest at most test locations during the 1993-94 growing season. Powdery mildew was not observed. Field disease notes for barley yellow dwarf and Septoria leaf blotch (Septoria tritici), were taken at locations where the incidence was severe enough to warrant. Barley yellow dwarf ratings were obtained by estimating the percent total necrotic flag leaves at the Columbia site (Table 6). Septoria leaf blotch was evaluated at the Columbia and Portageville locations (Table 6 and 10). Statistical Analyses and Interpretation The data collected at each soft red winter wheat location were analyzed as a four-replication, lattice design. Data collected from the hard red winter wheat locations were analyzed as a four- replication, randomized complete block design. If an observation was missing in one replication, the average of those observations in the remaining replications was used to approximate the missing observation. Fisher's least significant difference at the 0.05 probability level [LSD (p=0.05)] and coefficients of variation percentages (CV%) were calculated from analyses of variance by each location and across all locations. The LSD is used to compare the performance of two specific varieties at a time. If the mean of a variety exceeds that of another variety by more than the LSD, then the difference observed will be a true difference in 19 out of 20 instances under conditions similar to those of the test. Data collected on all traits measured during 1994 are presented in Tables 5 through 14 for the soft red wheats and in Tables 15 through 19 for the hard red wheats. Variety selection should be based on yield stability in a production environment over years and locations. Selection also should consider other characteristics such as test weight, plant height, heading date and disease resistance. Where these additional characteristics were not measured in a particular production environment, they can be evaluated from locations in which they were rated. Where a variety has been in the test for two or three years, combined analyses of the yield data over years are presented. Tables 14 and 19 rank all soft red winter and hard red winter wheats tested, respectively, according to their state-wide average. Overall rank can be very misleading. Differences in yield between any two varieties are considered significant or real only if that difference exceeds the LSD value given at the bottom of each column. Growers should be careful to make pair-wise comparisons of both the appropriate location(s) and state-wide averages before selecting one wheat variety over another for production in Missouri. 1994 Test Conditions Delayed emergence and poor fall tiller development from late planting dates and early cold weather were the primary concerns for winter wheat in Missouri during the 1993-94 cropping season, especially at the southeastern and southwestern locations. Adequate stands eventually did develop for most entries at all sites. The seed lots used to establish 'Coker 9543' and 'Coker 9803' expressed poor germination and produced poor stands at most test locations. Winter- kill and heaving problems did not develop to the degree observed in previous years. Winter survival averaged greater than 85 percent across all locations. Cool and drier than normal weather during late April and throughout May permitted spring tiller development. Dry weather and cool night-time temperatures occurred during the first 10 to 14 days after flowering. Conditions during this period were excellent for kernel development and grain fill. Conditions changed unusually hot and dry during the first two weeks of June, hastening maturity of the wheat particularly at the Columbia, Novelty and Trenton locations. Foliar diseases were not a major problem except at Portageville, which experienced substantial Septoria leaf blotch and some bacterial stripe. Overall yield of the 63 soft red winter wheats tested in 1994 was 65.5 bushels per acre, 21.4 bushels per acre more than the previous year and the highest average yield achieved in the last ten years of testing. Yields at the seven soft red winter wheat test locations varied from 51.4 bushels per acre at Mt. Vernon to 71.8 bushels per acre at Charleston (Table 14). 'Pioneer 2552' averaged 76.3 bushels per acre across all locations. Only 'FFR 525W' equaled the yield of Pioneer 2552 during the 1993-94 cropping season. The highest yielding public variety was 'Ernie', a recent release from the University of Missouri, averaging 72.5 bushels per acre. FFR 525W (63.2 bu/acre) and 'Pioneer 2548' (65.1 bu/acre)have been the highest yielding varieties tested for the past 2 and 3 years (Table 5), respectively. Although only two entries rank in the top yield group based on the 1994 state-wide average, the next five entries (Ernie, 'Wakefield', MO12258, 'BTX 105' and 'Madison') have yields statistically equal to those of FFR 525W, the second highest yielding variety (Table 14). Five additional entries ('AGRIPRO Hickory', 'Merschman Katie VII', 'Verne', 'AGRIPRO Clemens' and 'AGRIPRO Boone') of the 63 soft red winter wheat entries produced above average yields compared to the remaining 50 entries. Hard red winter wheat yields varied from 46.0 bushels per acre at Mt. Vernon to 77.4 bushels per acre at the Trenton site (Table 2). 'Karl 92' was highest yielding hard red winter wheat entry, averaging 65.3 bu/acre across three locations. Test weights among the soft red winter wheats were nearly 3 pounds per bushel heavier in 1994 as compared to test weights achieved in 1993 (see SR453, Winter Wheat - 1993 Missouri Crop Performance) and the heaviest test weights achieved since 1988. Location means varied from 55.8 pounds per bushel at Novelty (Table 7) to 60.4 pounds per bushel at Portageville (Table 10). Reductions in test weight at the Columbia, Novelty and Trenton sites are attributed to the hot and dry conditions that prevailed during the last two weeks of the grain fill period for these locations. Most entries had matured beyond the point of being substantially affected by these conditions at the Charleston, Portageville, Lamar and Mount Vernon locations. 'Coker 9474' produced the heaviest test weight at 60.8 pounds per bushel (Table 5). Eighteen other soft red winter wheat entries averaged test weights exceeding 58 pounds per bushel. Among the hard red winter wheats tested during 1994, 'Karl' averaged 61.1 pounds per bushels. Only Karl and Karl 92 produced test weights above the 60 pounds per bushel level needed for U.S. No. 1 hard red winter wheat. New Variety Descriptions Several entries previously tested as experimental lines have been or will be released as varieties for commercial production. Brief descriptions derived from variety release statements or other publicly available information are provided for informational purposes only. These descriptions do not imply endorsement or exclusion of any commercially available wheat varieties by the Missouri Agricultural Experiment Station. AGRIPRO Clemens Soft Red Winter Wheat ------------------------------------- AGRIPRO Clemens soft red winter wheat (PI583281, ABI 89-4580) is an awnless, mid-to-late season variety from AgriPro Seeds, Inc., with broad adaptation from the southern cornbelt to the northern mid-south regions of the United States. It was developed from the cross 'Pike / Coker 833' and was previously tested as 'ABI 89-4580' in regional trials. Plant Height of AGRIPRO Clemens is equal to that of 'Cardinal' wheat. Clemens is considered resistant to leaf rust, stem rust, Septoria leaf and glume blotches and powdery mildew. It is moderately resistant to soilborne mosaic virus, wheat spindle streak mosaic virus and Rhizoctonia root rot. Milling and baking quality is considered equal to or superior to that of 'Caldwell'. It also possesses the H3 gene, conferring resistance to biotypes GP, A and C of the Hessian fly. In Missouri during 1994, AGRIPRO Clemens produced above average yields combined with average test weights. Application for Plant Variety Protection has been made for AGRIPRO Clemens (PVP Cert. No. 9400167), with commercial sale of seed only as a class of certified seed. More detailed information on AGRIPRO Clemens is available from the company upon request. AGRIPRO Hickory Soft Red Winter Wheat ------------------------------------- AGRIPRO Hickory soft red winter wheat (PI573036, ABI 88-1903) was developed from the cross 'Hunter / Caldwell' by AgriPro Seeds, Inc. Hickory is an awnletted, early-to-mid season variety that is broadly adapted from southern Illinois to Maryland and from northern Louisiana to South Carolina. Winter survival has been excellent in its' area of adaptation, but Hickory may suffer injury in northern Missouri where more severe winter conditions frequently occur. Plant height is comparable to that of 'Wakefield'. AGRIPRO Hickory is considered moderately resistant to leaf rust, stem rust, powdery mildew, Septoria glume blotch, bacterial stripe and the soil virus complex caused by the soilborne mosaic and wheat spindle streak mosaic viruses. Milling and baking quality is good. In Missouri during 1994, AGRIPRO Hickory produced above average yields combined with average test weights. It is most likely best adapted to the Delta region of southeastern Missouri and on the more productive soil types west of the Ozarks in southwestern Missouri. Application for Plant Variety Protection has been made for AGRIPRO Hickory (PVP Cert. No. 9300291), with commercial sale of seed only as a class of certified seed. More detailed information on AGRIPRO Hickory is available from the company upon request. Ernie Soft Red Winter Wheat --------------------------- Ernie (MO12256) soft red winter wheat is a new variety developed by the Missouri Agricultural Experiment Station, from the cross 'Pike /3/ (MO9965, Stoddard / Blueboy // Stoddard / D1707)'. The variety is named after the late Dr. Ernest R. Sears, former USDA-ARS scientist at the University of Missouri-Columbia and world renown wheat cytogeneticist. Ernie is a short stature, awnletted, white-chaffed variety with excellent threshability and good shatter resistance. Maturity of Ernie is early, equal to that of Clark soft red winter wheat. Plant height of Ernie is approximately 1 and 6 inches shorter than Pioneer 2548 and Cardinal wheats, respectively. Ernie is moderately resistant to Septoria leaf blotch and powdery mildew. It has good tolerance to the barley yellow dwarf virus and appears to possess some field tolerance to scab. Ernie is susceptible to leaf rust, stem rust and the Hessian fly. Yield potential of Ernie is very good. Ernie was either the highest yielding or among the highest yielding entries at 4 of the 7 Missouri test locations in 1994 (Table 14). It ranked third highest in yield across all locations, only exceeded by Pioneer 2552 and FFR 525W wheats. Test weights were more than one pound per bushel heavier than the average of all entries (Table 5). Application for Plant Variety Protection of Ernie will be made under the Title V option by the Missouri Agricultural Experiment Station, with Ernie seed only available as Foundation or Certified classes of certified seed. Foundation seed should be available to certified seed growers in the fall of 1995. Grant Soft Red Winter Wheat --------------------------- Grant (PI572548, Pd811670A9-10-6-7-63) soft red winter wheat was developed by the Purdue University Agricultural Experiment Station from the cross 'Caldwell // Beau /Kavkaz'. It possesses the 1B/1R wheat/rye chromosome translocation obtained from the Kavkaz parent. Grant is an awnletted, white-chaffed variety, with overall plant type similar to that of Caldwell. Grant is approximately 2 inches shorter and heads 2 days later than Caldwell. Grant is considered resistant to powdery mildew, take-all, Septoria leaf and glume blotches, soilborne mosaic virus and wheat spindle streak mosaic virus. Grant possesses the Sr31 and Lr26 genes providing moderate stem and leaf rust resistance and the H5 gene, conferring resistance to biotypes GP, A, B, C, D and E of the Hessian fly. Grant was among the highest yielding wheats at Charleston (Table 9), but produced average to below average yields and below average test weights at the remaining test locations in Missouri during 1994 (Table 5 and 14). Grant is protect under the Title V option of the Federal Plant Variety Protect Act of 1970 (PVP Cert. No. 9300266), with seed of Grant only available as a class of certified seed. Hazen Soft Red Winter Wheat --------------------------- Hazen (AR26413A) Is an awned, white-chaffed, soft red winter wheat developed by the Arkansas Agricultural Experiment Station from the cross 'Doublecrop / Beau'. It is approximately 5 inches shorter in plant height and 2 days earlier in maturity than Cardinal soft red winter wheat. Hazen has excellent straw strength. Hazen is moderately resistant to leaf rust, soilborne mosaic virus and/or wheat spindle streak mosaic virus, and bacterial streak. Milling and baking quality for Hazen are acceptable. Hazen was among the highest yielding wheats at Charleston (Table 9). Hazen produced average yields combined with above average test weights across the remaining test locations in Missouri during 1994 (Table 5 and 14). The Arkansas Agricultural Experiment Station has not announced the availability of Hazen seed for commercial production in the fall of 1994. HybriTech Spencer Soft Red Winter Wheat --------------------------------------- HybriTech Spencer (PI547250, LB63) is a new proprietary soft red winter wheat developed by HybriTech Seed International, Inc., from the cross 'Arthur / Perdix // 2*Pd68243A3-9-1-80'. Spencer is an awned, white-chaffed variety possessing intermediate plant height, excellent straw strength and good winterhardiness. Heading date and maturity are approximately equal to that of Cardinal soft red winter wheat. Spencer is considered to have good tolerance to Septoria leaf blotch, Septoria glume blotch and tan spot. It is moderately resistant to soilborne mosaic virus and wheat spindle streak mosaic virus. Spencer is susceptible to the Hessian fly. Yields of Spencer were average across the seven test locations in Missouri during 1994. Test weights were excellent, over 3 pounds per bushel heavier than the average of all soft red winter wheats tested (Table 5). Spencer is protected under Title V of the Federal Plant Variety Protection Act of 1970, with seed only available as a class of certified seed. More detailed information on HybriTech Spencer is available from the company upon request. Ike Hard Red Winter Wheat ------------------------- Ike (PI574488, KS89H48-1) hard red winter wheat was developed at the Ft. Hays Branch Station of the Kansas Agricultural Experiment Station. It is an awned, white-chaffed selection from the cross 'Dular / Eagle // 2*Larned /Cheney /3/ Colt'. Ike is a tall semidwarf variety, averaging 3 to 4 more inches in plant height than Karl or Karl 92 hard red winter wheats and 1 to 2 inches shorter than Cardinal soft red winter wheat. Ike possesses excellent resistance to soilborne mosaic virus and wheat spindle streak mosaic virus, and is resistant to the Hessian fly. It is moderately resistant to stem rust and powdery mildew. Ike is moderately susceptible to leaf rust, Septoria leaf blotch and tan spot, and susceptible to the wheat streak mosaic virus. Milling and baking quality of Ike are very good. Yields of Ike in Missouri have been average to below average during 1994. Developed for production in western Kansas, Ike does not appear to be as well adapted as Karl or Karl 92 hard red winter wheats to the more humid conditions that exist in Missouri. Ike is protected by the Federal Plant Variety Protection Act of 1970 (PVP Cert. No. 9400149). Karl 92 Hard Red Winter Wheat ----------------------------- Karl 92 (PI564245, KS831374-142) is an awned, white-chaffed, mid- season, semidwarf hard red winter wheat developed by the Kansas Agricultural Experiment Station and the USDA Agricultural Research Service and released in 1992. Karl 92 is a single head-row selection from Karl hard red winter wheat. Karl 92 was selected for better leaf rust resistance and better green leaf duration. Karl 92 also heads about 1 day earlier than Karl. Karl 92 is considered moderately resistant to leaf rust, powdery mildew, tan spot, Septoria leaf blotch, Septoria glume blotch, bacterial leaf blight, soilborne mosaic virus and wheat spindle streak mosaic virus. It is susceptible to stem rust, wheat streak mosaic virus, greenbug and Hessian fly. Milling and baking quality are excellent. In 1994, Karl 92 was the highest yielding entry among five hard red winter wheats tested in Missouri (Tables 15 and 19). Karl 92 is protected under the Federal Plant Variety Protection Act of 1970 (PVP Cert. No. 9300124). Pioneer 2552 Soft Red Winter Wheat ---------------------------------- Pioneer 2552 (PI566924, WBC797E1) is a new proprietary soft red winter wheat developed by Pioneer Hi-Bred International, Inc. Pioneer 2552 is an awned, medium-to-late maturing variety, with short plant height and excellent straw strength. It has shown good adaptation across the southern cornbelt and the mid-south regions, including Missouri. Pioneer 2552 is moderately resistant to the predominant races of leaf rust in the northern soft wheat growing region, but has been more susceptible to races in the southeastern United States. It is considered resistance to powdery mildew, soil borne mosaic virus, wheat spindle streak mosaic virus, and possesses excellent tolerance to the common organisms that cause leaf blights in the soft wheat region. Pioneer does not recommend Pioneer 2552 in areas where wheat is prone to frequent winterkill. Pioneer 2552 has no known resistance to the Hessian fly. Pioneer 2552 averaged the highest yield across seven test locations in Missouri during 1994 (Tables 5 and 14). Test weights of Pioneer 2552 were 1.5 pounds per bushel heavier than the average of all soft red winter wheats tested. Pioneer 2552 is protected by the Federal Plant Variety Protection Act of 1970 (PVP Cert. No. 9300172). More detailed information on Pioneer 2552 is available from the company upon request. 1994 Missouri Winter Wheat Crop Projected Crop Statistics ------------------------- Missouri's 1994 wheat crop was harvested from an estimated 1.1 million acres, down 21 percent from the wheat acreage harvested in 1993 (Table 5). Substantial wheat acreage was not planted in the fall of 1993 due to very wet soil conditions and the delayed harvest of spring crops in many areas. The statewide average yield projected by the Missouri Agricultural Statistical Service is 44 bu/acre, up from 34 bu/acre and 40 bu/acre reported for 1991 and 1993, respectively. Projected district average yields ranged from a high of 48 bu/acre for the South-East district to a low of 38 bu/acre in the North-West district. Total projected production of the 1994 Missouri wheat crop is 48.4 million bushels, nearly 8 million bushels lower than 1993 production levels, representing a 14 percent decline. Electronic Accessibility of Data Results of the 1994 Missouri Winter Wheat Performance Tests are now available in two electronic forms. The winter wheat variety test data can be accessed on the Missouri Agricultural Bulletin Board (AgEBB) supported by the University of Missouri College of Agriculture, Food and Natural Resources. The telephone number for the AgEBB is (314) 882-8289. Baud rates up to 2,400 bps are presently supported. Select the "CROP PERFORMANCE TESTING" option from the main AgEBB menu. Call (314) 882-4827 to contact the AgEBB staff concerning questions or problems. For Internet users, the 1994 Missouri winter wheat test data can be obtained in the form of Excel 4.0 spreadsheets from the GrainGenes Gopher database located at Cornell University. The Internet host address for this Gopher database is 'greengenes.cit.cornell.edu port 70'. Select the "Grain files to download" menu option from the main GrainGenes menu and then select "Wheat Variety Tests". Missouri winter wheat variety test data from previous reports and reports from other states also are available. Acknowledgments Special Report No. 466 is a contribution of the Department of Agronomy, University of Missouri Agricultural Experiment Station. The Missouri Winter Wheat Performance Testing program is partially funded by Missouri wheat farmers and businessmen through a grant from the Missouri Seed Improvement Association and by fees from companies submitting varieties for evaluation. Special recognition goes to the following individuals who provided their time, land and additional resources in support of this program: Mr. Peter Brewer, Farmer Cooperator Trenton, Missouri Mr. Smith Deline, Farmer Cooperator Charleston, Missouri Mr. Jake Fisher, Superintendent Delta Research and Extension Center Portageville, Missouri Dr. Richard Crawford, Superintendent Southwest Research and Extension Center Mt. Vernon, Missouri Mr. John Poehlmann, Superintendent Bradford Agricultural Research Center Columbia, Missouri Mr. David Sheats, Farmer Cooperator Lamar, Missouri Mr. Randall Smoot, Superintendent Greenley Agricultural Research Center Novelty, Missouri The excellent cooperation and support provided by these individuals has been invaluable to this project. Special thanks is also extended to Mr. Bob Bellinghouse, Director, Missouri Agricultural Statistics Service, for supplying Missouri wheat production statistics and for help in direct mailing this publication out to Missouri wheat farmers. The authors would like to extend their appreciation to the following individuals for their contribution towards this report: Derrick Aulabaugh, John Bennett, Vera Reichlin, Kerry Strope, Sherry Teerlinck and Liming Wang. Authors: Kenneth D. Kephart, Assistant Professor and State Extension Agronomist-Small Grains; Anne L. McKendry, Assistant Professor and Soft Red Winter Wheat Breeder; David N. Tague, Senior Research Laboratory Technician; Dep. of Agronomy, University of Missouri, Columbia, MO; and James E. Berg, former Research Specialist, University of Missouri, presently Research Associate, Dep. of Plant, Soil and Environmental Sciences, Montana State University, Bozeman, MT.