Stock number: BGS 501
Locus name: Male sterile genetic 38
Locus symbol: msg38

Previous nomenclature and gene symbolization:

Male sterile jl = msg,,jl (2).

Monofactorial recessive (2).
Location is unknown.

Selfing - none (2).
Outcrossing - complete female fertility (2).
Stamens - anthers nearly normal in size, but without stomium, filament elongates (2).

A spontaneous mutant in Ingrid (CIho 10083) (2).

msg38.jl in Ingrid (MSS517) (1, 2).

msg38.jl in Ingrid (GSHO 2390); msg38.jl in Bowman (PI 483237)*7 (GSHO 2304).

1. Franckowiak, J.D., and E.A. Hockett. 1988. Identification of three new loci which control male sterility of barley. BGN 18:11-13.
2. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.

E.A. Hockett. 1991. BGN 20:105.

J.D. Franckowiak. 1997. BGN 26:427.

Stock number: BGS 502
Locus name: Male sterile genetic 39
Locus symbol: msg39

Previous nomenclature and gene symbolization:

Male sterile dm = msg,,dm (4).

Monofactorial recessive (1, 4).
Located in chromosome 6H, over 33.9 cM from the rob (orange lemma) locus (2).

Selfing - about 1% in Bozeman, Montana, USA and Elimäki, Finland (1, 4).
Outcrossing - complete female fertility (1).
Stamens - anthers are the same size as fertile sibs, stomium present, filament elongates (1, 4).

A spontaneous mutant in a Finnish six-rowed barley (P11, CIho 15836) (1, 4).

msg39.dm in P11 (MSS361) (3, 6); msg39.dn (MSS362) in a Finnish six-rowed barley (H31, CIho 15837) (1, 3, 6); msg39.ff (MSS407) in Sabarlis (CIho 15484) (3, 5).

msg39.dm in P11 (GSHO 2391); msg39.dm in Bowman (PI 483237)*7 (GSHO 2080).

1. Ahokas, H. 1976. Male sterile mutants of barley. III. Additional inaperturate mutants. BGN 6:4-6.
2. Franckowiak, J.D. 1991. Association of male sterility genes with a specific chromosome using multiple marker stocks. BGN 20:31-36.
3. Franckowiak, J.D., and E.A. Hockett. 1988. Identification of three new loci which control male sterility of barley. BGN 18:11-13.
4. Hockett, E.A. 1977. The genetic male sterile barley collection. BGN 7:97-100.
5. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
6. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1991. BGN 20:106.

J.D. Franckowiak. 1997. BGN 26:428.

Stock number: BGS 503
Locus name: Male sterile genetic 40
Locus symbol: msg40

Previous nomenclature and gene symbolization:

Male sterile ac = msg,,ac (3, 5).
Male sterile = ms-C (4).

Monofactorial recessive (3).
Located in chromosome 6H, over 27.8 cM from the rob (orange lemma) locus (1).

Selfing - none (3).
Outcrossing - less than 30% in crosses and in the Bowman backcross-derived line.
Stamens - anthers are smaller than those of fertile sibs, no stomium or filament elongation (3).

A spontaneous mutant in Conquest (CIho 11683) (4).

msg40.ac in Conquest (MSS095) (2, 3, 5).

msg40.ac in Conquest (GSHO 2393); msg40.ac in Bowman (PI 483237)*6 (GSHO 2081).

1. Franckowiak, J.D. 1991. Association of male sterility genes with a specific chromosome using multiple marker stocks. BGN 20:31-36.
2. Hockett, E.A. 1991. The genetic male sterile collection. Identification of eight new loci and allelism tests of 14 additional mutants. BGN 14:37-40.
3. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
4. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
5. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1991. BGN 20:107.

J.D. Franckowiak. 1997. BGN 26:429.

Stock number: BGS 504
Locus name: Male sterile genetic 41
Locus symbol: msg41

Previous nomenclature and gene symbolization:

Male sterile aj = msg,,aj (5).
Male sterile = 65msx166 (6).

Monofactorial recessive (5).
Location is unknown.

Selfing - none (3, 5).
Outcrossing - complete female fertility (5), but in the Bowman backcross-derived line seed set is less than 20% after crossing as a female parent.
Stamens - anthers rudimentary, no stomium or filament elongation (3, 5).

A spontaneous mutant in Betzes (PI 129430) (5).

msg41.aj in Betzes (MSS101) (4, 6, 7); msg41.dk in Betzes (MSS359) (1, 4, 7); msg41.do (MSS363) in Maris Baldric (PI 294512) (2, 4, 7); msg41.ef (MSS381) in Hector (CIho 15514), msg41.el (MSS387) in Sabarlis (CIho 15484), msg41.eq in Sel 12387Co (MSS392), msg41.fa (MSS402) in Midas (PI 343078), msg41.gl (MSS439) in Maris Mink (PI 467824), msg41.ij (MSS489) in Mazurka (PI 410868) (3, 4).

msg41.aj in Betzes (GSHO 2394); msg41.dk in Bowman (PI 483237)*6 (GSHO 2305).

1. Hockett, E.A. 1975. The genetic male sterile barley collection. BGN 5:84-86.
2. Hockett, E.A. 1977. The genetic male sterile barley collection. BGN 7:97-100.
3. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
4. Hockett, E.A. 1991. The genetic male sterile collection. Identification of eight new loci and allelism tests of 14 additional mutants. BGN 20:37-40.
5. Hockett, E.A., and R.F. Eslick. 1971. Genetic male sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
6. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
7. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1991. BGN 20:108.

J.D. Franckowiak. 1997. BGN 26:430.

Stock number: BGS 505
Locus name: Male sterile genetic 42
Locus symbol: msg42

Previous nomenclature and gene symbolization:

Male sterile db = msg,,db (2).
Male sterile = B68-N-109 (2).

Monofactorial recessive (2).
Located in chromosome 3H, over 18.9 cM from the alm (albino lemma) locus (1).

Selfing - none (2, 3).
Outcrossing - complete female fertility (2, 3).
Stamens - anthers rudimentary, no stomium or filament elongation (2, 3).

A spontaneous mutant in Betzes (PI 129430) (2).

msg42.db in Betzes (MSS350) (1, 4, 5); msg42.gt (MSS447) in Berac (PI 355136), msg42.hw in Sel 15025Co (MSS476), msg42.iy in Sv73608 (MSS504) (3, 4).

msg42.db in Betzes (GSHO 2395); msg42.db in Bowman (PI 483237)*6 (GSHO 1948).

1. Franckowiak, J.D. 1991. Association of male sterility genes with a specific chromosome using multiple marker stocks. BGN 20:31-36.
2. Hockett, E.A. 1975. The genetic male sterile barley collection. BGN 5:84-86.
3. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
4. Hockett, E.A. 1991. The genetic male sterile collection. Identification of eight new loci and allelism tests of 14 additional mutants. BGN 20:37-40.
5. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1991. BGN 20:109.

J.D. Franckowiak. 1997. BGN 26:431.

Stock number: BGS 506
Locus name: Male sterile genetic 43
Locus symbol: msg43

Previous nomenclature and gene symbolization:

Male sterile br = msg,,br (1).
Male sterile = B65msx38 (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - none (1).
Outcrossing - complete female fertility (1), but in the Bowman backcross-derived line seed set is less than 30% after crossing as a female parent.
Stamens - anthers rudimentary, no stomium or filament elongation (1).

A spontaneous mutant in Betzes (PI 129430) (1).

msg43.br in Betzes (MSS332) (1, 2, 3).

msg43.br in Betzes (GSHO 2396); msg43.br in Bowman (PI 483237)*7 (GSHO 2306).

1. Hockett, E.A. 1972. Coordinator's report on the genetic male sterile barley collection. BGN 2:139-144.
2. Hockett, E.A. 1991. The genetic male sterile collection. Identification of eight new loci and allelism tests of 14 additional mutants. BGN 20:37-40.
3. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1991. BGN 20:110.

J.D. Franckowiak. 1997. BGN 26:432.

Stock number: BGS 507
Locus name: Male sterile genetic 44
Locus symbol: msg44

Previous nomenclature and gene symbolization:

Male sterile cx = msg,,cx (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - none (1).
Outcrossing - complete female fertility (1).
Stamens - anthers rudimentary, no stomium or filament elongation (1).

A spontaneous mutant in selection HA6-33-02 (CIho 15835) (1).

msg44.cx in HA6-33-02 (MSS346) (1, 2, 3).

msg44.cx in HA6-33-02 (GSHO 2397); msg44.cx in Bowman (PI 483237)*7 (GSHO 2307).

1. Hockett, E.A. 1975. The genetic male sterile barley collection. BGN 5:84-86.
2. Hockett, E.A. 1991. The genetic male sterile collection. Identification of eight new loci and allelism tests of 14 additional mutants. BGN 20:37-40.
3. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1991. BGN 20:111.

J.D. Franckowiak. 1997. BGN 26:433.

Stock number: BGS 508
Locus name: Male sterile genetic 45
Locus symbol: msg45

Previous nomenclature and gene symbolization:

Male sterile dp = msg,,dp (1).
Male sterile = 76Y2 (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - none (1).
Outcrossing - complete female fertility (1).
Stamens - anthers rudimentary, no stomium or filament elongation (1).

A spontaneous mutant in selection RPB439-71 (CIho 15838) (1).

msg45.dp in RPB439-71 (MSS364) (1, 2, 3).

msg45.dp in RPB439-71(GSHO 2398); msg45.dp in Bowman (PI 483237)*4 (GSHO 2308).

1. Hockett, E.A. 1979. The genetic male sterile barley collection. BGN 9:124-128.
2. Hockett, E.A. 1991. The genetic male sterile collection. Identification of eight new loci and allelism tests of 14 additional mutants. BGN 20:37-40.
3. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1991. BGN 20:112.

J.D. Franckowiak. 1997. BGN 26:434.

Stock number: BGS 509
Locus name: Male sterile genetic 46
Locus symbol: msg46

Previous nomenclature and gene symbolization:

Male sterile ec = msg,,ec (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - none (1).
Outcrossing - complete female fertility (1).
Stamens - anthers smaller than those of fertile sibs, no stomium or filament elongation (1).

A spontaneous mutant in Hector (CIho 15514) (1).

msg46.ec in Hector (MSS378) (1, 2, 3).

msg46.ec in Hector (GSHO 2399); msg46.ec in Bowman (PI 483237)*4 (GSHO 2309).

1. Hockett, E.A. 1979. The genetic male sterile barley collection. BGN 9:124-128.
2. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
3. Hockett, E.A. 1991. The genetic male sterile collection. Identification of eight new loci and allelism tests of 14 additional mutants. BGN 20:37-40.

E.A. Hockett. 1991. BGN 20:113.

J.D. Franckowiak. 1997. BGN 26:435.

Stock number: BGS 510
Locus name: Male sterile genetic 47
Locus symbol: msg47

Previous nomenclature and gene symbolization:

Male sterile ep = msg,,ep (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - none (1).
Outcrossing - complete female fertility (1).
Stamens - anthers rudimentary, no stomium or filament elongation (1).

A spontaneous mutant in Sel 12384Co (1).

msg47.ep in Sel 12384Co (MSS391) (1, 2).

msg47.ep in Sel 12384Co (GSHO 2400); msg47.ep in Bowman (PI 483237)*5 (GSHO 2310).

1. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
2. Hockett, E.A. 1991. The genetic male sterile collection. Identification of eight new loci and allelism tests of 14 additional mutants. BGN 20:37-40.

E.A. Hockett. 1991. BGN 20:114.

J.D. Franckowiak. 1997. BGN 26:436.

Stock number: BGS 511
Locus name Reaction to Puccinia graminis 1 (wheat stem rust)
Locus symbol: Rpg1

Previous nomenclature and gene symbolization:

Resistance to Puccinia graminis tritici = T (8).
Resistance to Puccinia graminis Pers f sp tritici Eriks & E Henn = Rpg1a (4).

Stock number: BGS 512
Locus name Reaction to Puccinia graminis 2 (wheat stem rust)
Locus symbol: Rpg2

Previous nomenclature and gene symbolization:

Resistance to Puccinia graminis tritici = T2 (2).
Resistance to Puccinia graminis Pers f sp tritici Eriks & E Henn = Rpg2b (1).

Stock number: BGS 513
Locus name: Xantha seedling 2
Locus symbol: xnt2

Previous nomenclature and gene symbolization:

Xantha seedling b = x_b (1).
Yellow seedling b = x_b (2).

Monofactorial recessive (1).
Location is unknown.

Seedlings are yellow in color and do not develop green pigmentation. Since they do not survive beyond the two leaf stage, the xnt2.b gene must be maintained as a heterozygous stock (1).

A spontaneous mutant in Black Hulless (PI 24849) (1).

xnt2.b (x_b) in Black Hulless (Black Hulless II) (1).

xnt2.b in Black Hulless (GSHO 2).

1. Robertson, D.W. 1932. The effect of a lethal in the heterozygous condition on barley development. Colorado Agric. Exp. Stn. Tech. Bull. 1. 12 p.
2. Haus, T.E. 1988. BGS 513, Yellow seedling, x_b. BGN 18:82.

T.E. Haus 1988. BGN 18:82.

J.D. Franckowiak. 1997. BGN 26:440.

Stock number: BGS 515
Locus name: Reaction to Septoria passerinii 1
Locus symbol: Rsp1

Previous nomenclature and gene symbolization:

Resistance to Septoria passerinii Sacc = Sep (1, 2).

Monofactorial incomplete dominant (2).
Location is unknown (2).

The Rsp1.a gene conditions a high level of resistance to a single spore culture of Septoria passerinii isolated in Minnesota, USA. Infection occurs on F₁ seedlings, but is limited to a few lesions (2).

Natural occurrence in selection CIho 4249-2 (CIho 14300) (2).

Rsp1.a in CIho 14300.

Rsp1.a in CIho 14300 (GSHO 2510).

1. Moseman, J.G. 1972. Report on genes for resistance to pests. BGN 2:145-147.
2. Rasmusson, D.C., and W.E. Rogers. 1963. Inheritance of resistance to septoria in barley. Crop Sci. 3:161-163.

D.C. Rasmusson. 1988. BGN 18:84 as BGS 465.

J.D. Franckowiak. 1997. BGN 26:441.

Stock number: BGS 516
Locus name: Reaction to Septoria passerinii 2
Locus symbol: Rsp2

Previous nomenclature and gene symbolization:

Resistance to Septoria passerinii Sacc = Sep₂ (1, 2).

Monofactorial incomplete dominant (2).
Location is unknown, but linkage to the Rsp3 (reaction to Septoria passerinii 3) locus is reported (2).

The Rsp2.b gene conditions a high level of resistance to a single spore culture of Septoria passerinii isolated in Minnesota, USA. Pycnidia are observed in some, but not all lesions, on all F₁ plants (2).

Natural occurrence in accession CIho 4780 (PI 70837) (2).

Rsp2.b in PI 70837.

Rsp2.b in PI 70837 (GSHO 2511).

1. Moseman, J.G. 1972. Report on genes for resistance to pests. BGN 2:145-147.
2. Rasmusson, D.C., and W.E. Rogers. 1963. Inheritance of resistance to septoria in barley. Crop Sci. 3:161-163.

D.C. Rasmusson. 1988. BGN 18:85 as BGS 466.

J.D. Franckowiak. 1997. BGN 26:442.

Stock number: BGS 517
Locus name: Reaction to Septoria passerinii 3
Locus symbol: Rsp3

Previous nomenclature and gene symbolization:

Resistance to Septoria passerinii Sacc = Sep₃ (1, 2).

Monofactorial incomplete dominant (2).
Location is unknown (2), but linkage to the Rsp2 (reaction to Septoria passerinii 2) locus is reported (2)..

The Rsp3.c gene conditions a high level of resistance to a single spore culture of Septoria passerinii isolated in Minnesota, USA. Infection occurs on F₁ seedlings, but is limited to a few lesions (2).

Natural occurrence in selection II-51-43 from a Feebar/Kindred cross (CIho 10644) (2).

Rsp3.c in CIho 10644.

Rsp3.c in CIho 10644 (GSHO 2512).

1. Moseman, J.G. 1972. Report on genes for resistance to pests. BGN 2:145-147.
2. Rasmusson, D.C., and W.E. Rogers. 1963. Inheritance of resistance to septoria in barley. Crop Sci. 3:161-163.

D.C. Rasmusson. 1988. BGN 18:86 as BGS 467.

J.D. Franckowiak. 1997. BGN 26:443.

Stock number: BGS 518
Locus name: Semidwarf 1
Locus symbol: sdw1

Previous nomenclature and gene symbolization:

Denso dwarf = denso (4, 8).

Monofactorial recessive (4, 9), although some F₁'s tend to be intermediate in height compared to their parents (1, 5).
Location in chromosome 3HL (2, 6), probably proximal from the gsh2 (glossy sheath 2) locus.

Depending partially on environment, plants homozygous for the sdw1.a gene range from 10 to 30 cm shorter than normal sibs (1, 8, 10). Spike length is variable, but fully as long as in normal barley. The stock used for description of the sdw1.a gene, M21, has the short straw and long spike of the original 'Jotun Mutant' as well as a large culm diameter from its parent 'Vantage' (1, 10). The semidwarf mutant 'Diamant' is an allele at the sdw1 locus (4, 7). Alleles at the sdw1 locus are associated with semiprostrate juvenile growth (4, 8), delayed maturity (3, 8), and reduced malt quality (3, 9).

An X-ray induced mutant in the Norwegian cultivar Jotun (PI 467357) isolated as Jotun 22 by Knut Mikaelsen (1, 5).

sdw1.a in Jotun (66/86, GSHO 1414) (10); sdw1.c (denso) in an unknown cultivar (4); sdw1.d (Diamant) in Valticky (4); sdw1.e (Risø 9265) in Bomi (PI 43371) (4).

sdw1.a in M21 (CIho 15481, GSHO 2513) from the cross Jotun Mutant/Kindred// Vantage (9); sdw1.d in Triumph (CIho 11612, GSHO 2465); sdw1.a in Bowman (PI 483237)*7 (GSHO 1978); sdw1.d from Triumph in Bowman*4 (GSHO 1979).

1. Ali, M.A.M., O. Okiror, and D.C. Rasmusson. 1978. Performance of semidwarf barley. Crop Sci. 18:418-422.
2. Barau, U.M., K.J. Chambers, W.T.B. Thomas, C.A. Hackett, V. Lea, P. Jack, B.P. Forster, R. Waugh, and W. Powell. 1993. Molecular mapping of genes determining height, time to heading, and growth habit in barley (Hordeum vulgare). Genome 36:1080-1087.
3. Foster, A.E., and A.P. Thompson. 1987. Effects of a semidwarf gene from Jotun on agronomic and quality traits of barley. p. 979-982. In S. Yasuda and T. Konishi (eds.) Barley Genetics V., Proc. Fifth Int. Barley Genet. Symp., Okayama, 1986. Sanyo Press Co., Okayama.
4. Haahr, V., and D. von Wettstein. 1976. Studies of an induced, high-yielding dwarf-mutant of spring barley. p. 215-218. In H. Gaul (ed.) Barley Genetics III., Proc. Third Int. Barley Genet. Symp., Garching, 1975. Verlag Karl Thiemig, München.
5. Lambert, J.W., and M. Shafi. 1959. Inheritance and heritability of height in three barley crosses. Barley Newsl. 3:7-8. (Abstr.)
6. Laurie, D.A., N. Pratchett, C. Romero, E. Simpson, and J.W. Snape. 1993. Assignment of the denso dwarfing gene to the long arm of chromosome 3 (3H) of barley by use of RFLP markers. Plant Breed. 111:198-203.
7. Mickelson, H.R., and D.C. Rasmusson. 1994. Genes for short stature in barley. Crop Sci. 34:1180-1183.
8. Powell, W., P.D.J. Caligari, W.T.B. Thomas, and J.L. Jinks. 1985. The effects of major genes on quantitatively varying characters in barley. 2. The denso and day length response loci. Heredity 54:349-352.
9. Powell, W., P.D.J. Caligari, W.T.B. Thomas, and J.L. Jinks. 1991. The effects of major genes on quantitatively varying characters in barley. 4. The GPert and denso loci and quality characters. Heredity 66:381-389.
10. Rasmusson, D.C., E.E. Banttari, and J.W. Lambert. 1973. Registration of M21 and M22 semidwarf barley. Crop Sci. 13:777.

D.C. Rasmusson. 1988. BGN 18:87 as BGS 468.

J.D. Franckowiak. 1997. BGN 26:444-445.

Stock number: BGS 519
Locus name: Many-noded dwarf 1
Locus symbol: mnd1

Previous nomenclature and gene symbolization:

Many noded dwarf = m (2, 4).

Monofactorial recessive (2, 3, 4).
Location is unknown.

When grown in the field, plants are about 1/2 normal height with numerous tillers. Each tiller has 12 to 20 small leaves and a spike of 1/2 normal size or less. Additional tillers with 4 to 7 leaves may develop at the uppermost tiller nodes, but these secondary and tertiary culm branches develop late and have very small spikes (2). When grown in the greenhouse, plants are taller and later than normal sibs and each tiller may have 15 to 20 nodes. The rate of primordium initiation is normal, but their expansion into leaves is much more rapid (1).

A spontaneous mutant in Mesa (CIho 2328) (2).

mnd1.a in Mesa (2).

mnd1.a in Mesa (GSHO 253); mnd1.a in Bowman (PI 483237)*9 (GSHO 2038).

1. Cooper, J.P. (unpublished).
2. Harlan, H.V., and M.N. Pope. 1922. Many-noded dwarf barley. J. Hered. 13:269-273.
3. Hor, K.S. 1922. A new variety of barley with striking characteristics. Science 55:378.
4. Hor, K.S. 1924. Interrelations of genetic factors in barley. Genetics 9:151-180.

J.P. Cooper and J.D. Franckowiak. 1997. BGN 26:446.

Stock number: BGS 520
Locus name: Male sterile genetic 48
Locus symbol: msg48

Previous nomenclature and gene symbolization:

Male sterile genetic jt = msg,,jt and Msg76-5 (3).

Monofactorial recessive (2, 3).
Located in chromosome 2H, more than 31.9 cM from the vrs1 (six-rowed spike 1) locus (1).

Selfing - 5% (2), but it may be near 50% in certain environments.
Outcrossing - Complete female fertility (2).
Stamens - anthers slightly smaller than fertile sib with filament elongation, but no stomium (2).

A spontaneous mutant in Simba (PI 584816, NGB 1505) (2).

msg48.jt in Simba (MSS525) (2, 3).

msg48.jt in Simba (GSHO 2401); msg48.jt in Bowman (PI 483237)*4 (GSHO 1925).

1. Franckowiak, J.D. 1991. Association of male sterility genes with a specific chromosome using multiple marker stocks. BGN 20:31-36.
2. Franckowiak, J.D., and E.A. Hockett. 1992. Allelism tests for the genetic male sterile msg,,jt. BGN 21:23-24.
3. Hockett, E. A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.

J.D. Franckowiak and E.A. Hockett. 1997. BGN 26:447.

Stock number: BGS 521
Locus name: Mottled leaf 1
Locus symbol: mtt1

Previous nomenclature and gene symbolization:

Mottled leaves = mt (3).
Mottled leaves 3 = mt₃ (2).

Monofactorial recessive (3).
Located in chromosome 1HS [5S] (2), based on linkage drag with the sls1.a (small lateral spikelet) gene (1).

Seedlings have two to four clearly marked white bands across or partially across the leaves. Necrosis of the white region occurs later and the leaf is constricted in these regions. Temperatures at germination over 20 C reduce the extent of mottling. Plant mortality is low at Edmonton, Alberta, Canada, where temperatures are low, but nearly 100% at St. Paul, Minnesota, USA (2).

An induced mutant in Montcalm (CIho 7149) (2).

mtt1.a (mt) in Montcalm (Alb Acc 284) (2).

mtt1.a in Montcalm (OUM070, GSHO 622); mtt1.a in Bowman (PI 483237)*4 (GSHO 2036).

1. Franckowiak, J.D. 1995. Notes on linkage drag in Bowman backcross derived lines of spring barley. BGN 24:63-70.
2. Kasha, K.J., and G.W.R. Walker. 1960. Several recent barley mutants and their linkages. Can. J. Genet. Cytol. 2:397-415.
3. Walker, G.W.R., K. Kasha, and R.A. Miller. 1958. Recombination studies in barley. Proc. Genet. Soc. Can. 3:41-43.

J.D. Franckowiak. 1997. BGN 26:448.

Stock number: BGS 522
Locus name: Eceriferum-yi
Locus symbol: cer-yi

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the leaf blade appears absent (wax code ++ ++ -) (2).

An X-ray induced mutant in Foma (CIho 11333) (1).

cer-yi.254 in Foma, -yi.956 in Bonus (PI 189763) (2); cer-yi.1213 in Kristina, -yi.1353 in Bonus (3).

cer-yi.254 in Foma (GSHO 1542); cer-yi.254 in Bowman (PI 483237)*3 (GSHO 2213).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.
3. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:449.

Stock number: BGS 523
Locus name: Eceriferum-yj
Locus symbol: cer-yj

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the leaf blade appears absent on the upper three leaves (wax code ++ ++ -) (2). Plants of the original stock are semidwarf (about 3/4 normal height), but those of the Bowman backcross-derived line are not.

A neutron induced mutant in Bonus (PI 189763) (1).

cer-yj.667 in Bonus, -yj.1217 in Kristina (2); cer-yj.937 in Bonus (3); cer-yj.1150 in Kristina, -yj.1811 in Bonus (4).

cer-yj.667 in Bonus (GSHO 1543); cer-yj.667 in Bowman (PI 483237)*3 (GSHO 2214).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.
3. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
4. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:450.

Stock number: BGS 524
Locus name: Eceriferum-yk
Locus symbol: cer-yk

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike, leaf sheath, and stem apppears reduced (wax code + + ++) (2). The original stock has narrow leaves, a lax spike, and extra wax coating on the kernels. Plants of the Bowman backcross-derived line seem to lack surface wax on the nodes (glossy node).

A neutron induced mutant in Bonus (PI 189763) (1).

cer-yk.627 in Bonus (2).

cer-yk.627 in Bonus (GSHO 1544); cer-yk.627 in Bowman (PI 483237)*4 (GSHO 2215).

1. Lundqvist, U. (unpublished).
2. Lundqvist. U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:451.

Stock number: BGS 525
Locus name: Eceriferum-yl
Locus symbol: cer-yl

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike, the leaf sheath, and the stem appears absent (wax code - - ++). The original mutant stock produces weak plants (about 3/4 normal height) with a high level of sterility (2). Plants in the Bowman backcross-derived line are semidwarf with waxy nodes, coiled peduncles, globose shaped seeds, and poorly attached hull. Reduced seed set can be caused by adverse environmental conditions.

A propane disulfonic acid diethyl ester induced mutant in Bonus (PI 189763) (1).

cer-yl.187, -yl.188 in Bonus (2); cer-yl.407 in Foma (CIho 11333), -yl.821 in Bonus (3).
Mutant used for description and seed stocks:
cer-yl.187 in Bonus (GSHO 1545); cer-yl.187 in Bowman (PI 483237)*6 (GSHO 2216).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.
3. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:452.

Stock number: BGS 526
Locus name: Eceriferum-ym
Locus symbol: cer-ym

Previous nomenclature and gene symbolization:

None.
Inheritance:
Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike, the leaf sheath and stem, and the leaf blade appears absent (wax code - - -) (2). Plants are weak and semidwarf (2/3 normal height) and heading is delayed. In the Bowman backcross-derived line, plants are weak, culms have a kinked peduncle, nodes are wax covered, and seeds are small, globe shaped, and nearly hulless.

An ethylene oxide induced mutant in Bonus (PI 189763) (1).

cer-ym.130, -ym.753 in Bonus (2).

cer-ym.753 in Bonus (GSHO 1546); cer-ym.753 in Bowman (PI 483237)*5 (GSHO 2217).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:453.

Stock number: BGS 527
Locus name: Eceriferum-yn
Locus symbol: cer-yn

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike, leaf sheath, and stem is reduced (wax code
+ + ++) (2). In the Bowman backcross-derived line, the wax coating is only slightly reduced, but nodes are glossy.

An ethylene imine induced mutant in Kristina (1).

cer-yn.1112 in Kristina (2).

cer-yn.1112 in Kristina (GSHO 1547); cer-yn.1112 in Bowman (PI 483237)*2 (GSHO 2218).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:454.

Stock number: BGS 528
Locus name: Eceriferum-yo
Locus symbol: cer-yo

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (1).
Location is unknown.

Surface wax coating on the leaf blade is reduced (wax code ++ ++ +) (2). The original stock in Bonus has a high level of sterility. Seedlings of the original stock and the Bowman backcross-derived line have slightly pale green color.

An ethyl methanesulfonate induced mutant in Bonus (PI 189763) (1).

cer-yo.647 in Bonus (2).

cer-yo.647 in Bonus (GSHO 1548); cer-yo.647 in Bowman (PI 483237)*3 (GSHO 2219).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:455.

Stock number: BGS 529
Locus name: Eceriferum-yp
Locus symbol: cer-yp

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the leaf blade is reduced (wax code ++ ++ +) (2). Leaves have a pale green color similar that of some chlorina mutants.

A neutron induced mutant in Bonus (PI 189763) (1).

cer-yp.949, -yp.955 in Bonus (2).

cer-yp.949 in Bonus (GSHO 1549); cer-yp.949 in Bowman (PI 483237)*2 (GSHO 2220).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:456.

Stock number: BGS 530
Locus name: Eceriferum-yq
Locus symbol: cer-yq

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the leaf blade appears absent (wax code ++ ++ -) (2). In the Bowman backcross-derived line, the wax coating is reduced only on upper leaf blades (wax code ++ ++ +).

An ethyl methanesulfonate induced mutant in Kristina (1).

cer-yq.1246 in Kristina (2).

cer-yq.1246 in Kristina (GSHO 1550); cer-yq.1246 in Bowman (PI 483237)*3 (GSHO 2221).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:457.

Stock number: BGS 531
Locus name: Eceriferum-yr
Locus symbol: cer-yr

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike appears greatly reduced or absent, and surface wax coating on the leaf sheath and stem is reduced (wax code -/+ + ++) (2). Nodes are glossy and appear to lack surface wax.

An ethylene imine induced mutant in Foma (CIho 11333) (1).

cer-yr.492 in Foma, -yr.1418 in Bonus (PI 189763) (2).

cer-yr.492 in Foma (GSHO 1551).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:458.

Stock number: BGS 532
Locus name: Eceriferum-ys
Locus symbol: cer-ys

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the leaf blade appears absent (wax code ++ ++ - ) (2). The original stock has a lax spike.

An ethylene imine induced mutant in Bonus (PI 189763) (1).

cer-ys.680, -ys.1352 in Bonus (2); cer-ys.1182, -ys.1247 in Kristina (3); cer-ys.1826 in Bonus (4).

cer-ys.680 in Bonus (GSHO 1552); cer-ys.680 in Bowman (PI 483237)*3 (GSHO 2222).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
3. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15.89-93.
4. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:459.

Stock number: BGS 533
Locus name: Eceriferum-yt
Locus symbol: cer-yt

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike appears absent (wax code - ++ ++) (2). In the Bowman backcrossed-derived line, nodes appear to lack surface wax.

An X-ray induced mutant in Bonus (PI 189763) (1).

cer-yt.758, -yt.938, -yt.1333 in Bonus (2); cer-yt.1127, -yt.1304 in Kristina, -yt.1427, -yt.1458, -yt.1493, -yt.1766, -yt.1822 in Bonus (3); cer-yt.1379 in Bonus (4).

cer-yt.938 in Bonus (GSHO 1553); cer-yt.938 in Bowman (PI 483237)*4 (GSHO 2223).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
3. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
4. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:460.

Stock number: BGS 534
Locus name: Eceriferum-yu
Locus symbol: cer-yu

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the leaf blade appears absent only on upper leaves (wax code ++ ++ -) (2). Mutant plants have slightly coiled awns. The Bowman backcross-derived line is weak in the field and has short leaves, but these traits are not as obvious in greenhouse grown plants.

A neutron induced mutant in Bonus (PI 189763) (1).

cer-yu.158, -yu.159, -yu.174, -yu.992, -yu.1799 in Bonus, -yu.1168, -yu.1170 in Kristina (2).
Mutant used for description and seed stocks:
cer-yu.159 in Bonus (GSHO 1554); cer-yu.159 in Bowman (PI 483237)*3 (GSHO 2224).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:461.

Stock number: BGS 535
Locus name: Eceriferum-yx
Locus symbol: cer-yx

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike, the leaf sheath, and stem appears reduced (wax code + + ++) (2).

A neutron induced mutant in Foma (CIho 11333) (1).

cer-yx.421 in Foma, -yx.1226 in Kristina (2); cer-yx.1092, -yx.1094 in Kristina (3).

cer-yx.421 in Foma (GSHO 1555); cer-yx.421 in Bowman (PI 483237)*2 (GSHO 2225).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
3. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:462.

Stock number: BGS 536
Locus name: Eceriferum-yy
Locus symbol: Cer-yy

Previous nomenclature and gene symbolization:

Glaucous head = ge (9).
Glossy spike 1 = Gle1 (8).

Monofactorial dominant (5, 8).
Located in chromosome 1HS [5S] (8), about 4.2 cM distal from the Hor2 (hordein 2, HorB) locus (1, 8).

Surface wax coating on the spike appears absent (wax code - ++ ++) (5). Concerning the chemical epicuticular wax composition, the Cer-yy mutant in Bonus determines waxes lacking -hydroxy- -diketones and having relatively increased proportions of primary alcohols. The distribution of the lipid classes is similar to that of wild type plants (7).

The Cer-yy gene or an allele is present in many Hordeum vulgare subsp spontaneum accessions and in a few Hordeum vulgare subsp vulgare cultivars; an ethyl methanesulfonate induced mutant in Bonus (PI 189763) (2).

Gle1.a in Cristal (3, 4, 8, 10); Cer-yy.437 in Foma (CIho 11333), -yy.649, -yy.650, -yy.849, -yy.898, -yy.919, -yy.920, -yy.923, -yy.929, -yy.935, -yy.941, -yy.966, -yy.968, -yy.975, -yy.982, -yy.983, -yy.984 in Bonus (5); Cer-yy.974, -yy.1717 in Bonus (6).

Cer-yy.849 in Bonus (GSHO 1556); Gle1.a in Estate (CIho 3410, GSHO 1316); Cer-yy.849 in Bowman (PI 483237)*6 (GSHO 2029); Gle1.a from Estate in Bowman*7 (GSHO 2030).

1. Jensen, J. 1987. Coordinator's report: Chromosome 5. BGN 17:111-113.
2. Lundqvist, U. (unpublished).
3. Lundqvist, U. 1991. Coordinator's report: Eceriferum genes. BGN 20:84.
4. Lundqvist, U. 1992. Coordinator's report: Eceriferum genes. BGN 21:99-101.
5. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
6. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.
7. Lundqvist, U., and P. von Wettstein-Knowles. 1982. Dominant mutations at Cer-yy change barley spike wax into leaf blade wax. Carlsberg Res. Commun. 47:29-43.
8. Netsvetaev, V.P., and A.A. Sozinov. 1982. Linkage studies of genes Gle1 and HrdF in barley chromosome 5. BGN 12:13-18.
9. Robertson, D.W., G.A. Wiebe, and R.G. Shands. 1947. A summary of linkage studies in barley: Supplement I, 1940-1946. J. Am. Soc. Agron. 39:464-473.
10. Wettstein-Knowles, P. von. 1991. New alleles of Cer-yy and cer-b. BGN 20:66-68.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:463.

Stock number: BGS 537
Locus name: Eceriferum-yz
Locus symbol: cer-yz

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike, leaf sheath, and stem appears slightly reduced (wax code + + ++) (2).

A sodium azide induced mutant in Bonus (PI 189763) (1).

cer-yz.1452, -yz.1453 in Bonus (2).

cer-yz.1452 in Bonus (GSHO 1557); cer-yz.1452 in Bowman (PI 483237)*2 (GSHO 2226).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:464.

Stock number: BGS 538
Locus name: Eceriferum-xa
Locus symbol: cer-xa

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the leaf blade appears absent only on upper leaves (wax code ++ ++ -) (2).

A gamma-ray induced mutant in Foma (CIho 11333) (1).

cer-xa.585 in Foma, -xa.838 in Bonus (PI 189763), -xa.1113 in Kristina (2).

cer-xa.585 in Foma (GSHO 1558); cer-xa.585 in Bowman (PI 483237)*2 (GSHO 2227).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:465.

Stock number: BGS 539
Locus name: Eceriferum-xb
Locus symbol: cer-xb

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike appears absent (wax code - ++ ++) (2). In the Bowman backcross-derived line, the peduncle appears to lack surface wax.

An isopropyl methanesulfonate induced mutant in Bonus (PI 189763) (1).

cer-xb.943 in Bonus, -xb.1166 in Kristina (2).

cer-xb.943 in Bonus (GSHO 1559); cer-xb.943 in Bowman (PI 483237)*4 (GSHO 2228).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:466.

Stock number: BGS 540
Locus name: Eceriferum-xc
Locus symbol: cer-xc

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike, leaf sheath, and stem is reduced (wax code
+ + ++) (2).

A sodium azide induced mutant in Bonus (PI 189763) (1).

cer-xc.1371, -xc.1457 in Bonus (2).

cer-xc.1371 in Bonus (GSHO 1560); cer-xc.1371 in Bowman (PI 483237)*2 (GSHO 2229).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:467.

Stock number: BGS 541
Locus name: Eceriferum-xd
Locus symbol: cer-xd

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2).
Location is unknown.

Surface wax coating on the spike, leaf sheath, and stem is reduced (wax code
+ + ++) (2). The Bowman backcross-derived line seems to have extra surface wax on the nodes.

A sodium azide induced mutant in Bonus (PI 189763) (1).

cer-xd.1455, -xd.1492 in Bonus (2).

cer-xd.1455 in Bonus (GSHO 1561); cer-xd.1455 in Bowman (PI 483237)*2 (GSHO 2230).

1. Lundqvist, U. (unpublished).
2. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:468.

Stock number: BGS 543
Locus name: Intermedium spike-f
Locus symbol: int-f

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (1, 2).
Location is unknown.

The spike appears six-rowed, but the lateral spikelets are much smaller, less than half the size of the central spikelets. Lateral spikelets are pointed and often have short awns. Seed set occurs in the lateral spikelets in the upper third of the spike. The base of the spike has shortened rachis internodes and appears erectoides-like (3).

An ethyl methanesulfonate induced mutant in Foma (CIho 11333) (3).

int-f.19 in Foma (2, 3).

int-f.19 in Foma (GSHO 1767); int-f.19 in Bowman (PI 483237)*3 (GSHO 2270).

1. Gustafsson, Å., A. Hagberg, U. Lundqvist, and G. Persson. 1969. A proposed system of symbols for the collection of barley mutants at Svalöv. Hereditas 62:409-414.
2. Gustafsson, Å., and U. Lundqvist. 1980. Hexastichon and intermedium mutants in barley. Hereditas 92:229-236.
3. Lundqvist, U., and A. Lundqvist. 1988. Induced intermedium mutants in barley: origin, morphology and inheritance. Hereditas 108:13-26.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:469.

Stock number: BGS 544
Locus name: Intermedium spike-h
Locus symbol: int-h

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (1).
Location is unknown.

Lateral spikelets are enlarged and have an inconspicuously pointed apex, but they do not set seed. Induced mutants show early heading and have an elongated basal rachis internode. The spike appears lax with shortened rachis internodes at the base (1). The Bowman backcross-derived line is slightly shorter (5/6 normal) and produces extra spikelets (up to five fertile ones) at several rachis nodes in the lower half of the spike. Fusion of some spikelets results in double and occasionally triple kernels.

A neutron induced mutant in Kristina (1, 2).

int-h.42, -h.43, -h.44 in Kristina (1, 2); int-h.83 in Bonus (PI 189763).

int-h.42 in Kristina (GSHO 1768); int-h.42 in Bowman (PI 483237)*5 (GSHO 2271).

1. Gustafsson, Å., and U. Lundqvist. 1980. Hexastichon and intermedium mutants in barley. Hereditas 92:229-236.
2. Lundqvist, U., and A. Lundqvist. 1988. Induced intermedium mutants in barley: origin, morphology and inheritance. Hereditas 108:13-26.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:470.

Stock number: BGS 545
Locus name: Intermedium spike-i
Locus symbol: int-i

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (1).
Location is unknown.

Lateral spikelets are enlarged and slightly pointed at the apex, but they fail to set seed. The tip of the spike may have very short rachis internodes and appears very dense or fasciated (1). The Bowman backcross-derived line exhibits reduced plant height (5/6 normal), reduced spike length, and a distinct elongation of the basal rachis internode.

An ethylene imine induced mutant in Kristina (2).

int-i.39 in Kristina (1, 2).

int-i.39 in Kristina (GSHO 1769); int-i.39 in Bowman (PI 483237)*4 (GSHO 2272).

1. Gustafsson, Å., and U. Lundqvist. 1980. Hexastichon and intermedium mutants in barley. Hereditas 92:229-236.
2. Lundqvist, U., and A. Lundqvist. 1988. Induced intermedium mutants in barley: origin, morphology and inheritance. Hereditas 108:13-26.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:471.

Stock number: BGS 546
Locus name: Intermedium spike-k
Locus symbol: int-k

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (1).
Location is unknown.

The spike is short and dense in the original mutant. Lateral spikelets are enlarged and the apex is pointed, and they occasionally have a short awn. Seed set does not occur in lateral spikelets and central spikelets are semi-sterile (1). Plants of the original stock have a dense coating of surface waxes. In the Bowman backcross-derived line, plants are small and weak (about 1/2 normal height) and have short spikes (1/2 normal), reduced awn length (3/4 normal), and very poor seed set. Awns of plants in the derived line are semi-rough, but F₁ hybrids with Bowman have semismooth awns.

An ethyl methanesulfonate induced mutant in Kristina (1).

int-k.47 in Kristina (1).

int-k.47 in Kristina (GSHO 1770); int-k.47 in Bowman (PI 483237)*6.

1. Lundqvist, U., and A. Lundqvist. 1988. Induced intermedium mutants in barley: origin, morphology and inheritance. Hereditas 108:13-26.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:472.

Stock number: BGS 547
Locus name: Intermedium spike-m
Locus symbol: int-m

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (3).
Location is unknown.

The spike is very short and has irregular rachis internode lengths. Lateral spikelets are enlarged and pointed, but they do not set seed. Spikelet density at the base of the spike is increased. Rachis internodes at the tip of the spike are very short, and the spike appears to have two or three fused or fasciated terminal spikelets. Tillering of int-m plants is increased and heading is slightly earlier (3).

A sodium azide induced mutant in Bonus (PI 189763) (2).

int-m.85 in Bonus (3); int-m.la (GSHO 1773) in Lamont (PI 512036) (1).

int-m.85 in Bonus (GSHO 1772); int-m.85 in Bowman (PI 483237)*5 (GSHO 2273); int-m.la in Bowman (PI 483237)*4 (GSHO 2274).

1. Franckowiak, J.D. (unpublished).
2. Lundqvist, U. (unpublished).
3. Lundqvist, U., and A. Lundqvist. 1988. Induced intermedium mutants in barley: origin, morphology and inheritance. Hereditas 108:13-26.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:473.

Stock number: BGS 548
Locus name: Angustifolium-b
Locus symbol: Fol-b

Previous nomenclature and gene symbolization:

Angustifolia = Ang (2).

Monofactorial incomplete dominant (1).
Location is unknown, but a possible association with chromosome 1H [5] based on trisomic analysis (2).

Homozygous plants are very weak with short thread-like leaves and often die at the 3 to 4 leaf stage. Heterozygotes have narrow leaves, small spikes and kernels, but vigor is good and fertility is normal (1).

A spontaneous mutant in an unknown cultivar (1).

Fol-b.2 in an unknown cultivar (1).

Fol-b.2 in an unknown cultivar (GSHO 17).

1. Tsuchiya, T., and R.L. Haines. 1975. Trisomic analysis of nine mutant genes in barley. BGN 5:67-69.
2. Tsuchiya, T., S.D. Tanksley, and R. Ward. 1976. Trisomic analysis of three mutant genes in barley. BGN 6:95.

J.D. Franckowiak. 1997. BGN 26:474.

Stock number: BGS 549
Locus name: Long glume awn
Locus symbol: Lga

Previous nomenclature and gene symbolization:

Long glume awn = L (2, 3).
Long outer glume = Log (1); this gene symbol is recommended by Helgason (1), but the symbol is also used for the eog (elongated outer glume) gene (4).

Monofactorial dominant (1, 2).
Location is unknown.

In spikelets of plants with the dominant allele (Lga1.a), the glume (outer) plus its awn is much longer than the kernel. Heterozygotes have a glume awn of intermediate length. In the short glume awn phenotype (lga1.b), the glume length and width are unaltered, but the glume plus its awn are about the same length as the kernel. Tsuchiya (4) reports that the glume awn length is 4 to 6 mm for the short type and 11 to 13 mm for the long type. Recessive alleles at the eog locus (see BGS 57) produce an array of glume sizes, some of which have glumes phenotypically similar to those associated with the dominant allele at the Lga locus. However, large glumes controlled by eog alleles are wider than normal and show a recessive inheritance pattern.

The short glume awn appears to be the mutant phenotype and occurs commonly in two-rowed cultivars of Occidental origin.

Lga1.a in Guy Mayle (CIho 620) (2, 3); lga1.b in Canadian Thorpe (PI 2710) (2, 3).

Lga1.a in Guy Mayle (GSHO 835); lga1.b in Canadian Thorpe (GSHO 834); lga1.b in Bowman (PI 483237) from Hector (CIho 15514); Lga1.a from Akashinriki (OUJ659, PI 467400) mutant OUM113 in Bowman (PI 483237)*8 (GSHO 2147).

1. Helgason, S.B. 1964. Inheritance of glume characteristics in some barley crosses. p. 181-185. In S. Broekhuizen, G. Dantuma, H. Lamberts, and W. Lange (eds.) Barley Genetics I. Proc. First Int. Barley Genet. Symp., Wageningen, 1963. Centre for Agricultural Publications and Documentation, Wageningen.
2. Neatby, K.W. 1926. Inheritance of quantitative and other characters in a barley cross. Sci. Agric. 7:77-84.
3. Neatby, K.W. 1929. An analysis of the inheritances of quantitative characters and linkage in barley. Sci. Agric. 9:701-718.
4. Tsuchiya, T. 1974. Preliminary results on genetic studies of outer glume characters in barley. BGN 4:76-78.

J.D. Franckowiak and U. Lundqvist. 1997. BGN 26:475.

Stock number: BGS 550
Locus name: Breviaristatum-b
Locus symbol: ari-b

Previous nomenclature and gene symbolization:

Breviaristatum-11 = ari-11 (1, 2).

Monofactorial recessive (2).
Location is unknown.

Plants are semidwarf with reduced viability and fertility; leaves are broad, but shortened and lighter green in color; and awns are about 3/4 normal length. Growth is markedly retarded during warm seasons and relatively good during cool seasons (2). Development of reproductive organs is delayed and sterility causes a stock maintenance problem.

A gamma-ray induced mutant in Bonus (PI 189763) (2).

ari-b.11, -b.13, -b.17, -b.19, -b.29 in Bonus, -b.175, -b.226, -b.238 in Foma (CIho 11333) (2).

ari-b.11 in Bonus (GSHO 1649); ari-b.11 in Bowman (PI 483237)*3 (GSHO 2157).

1. Gustafsson, Å., A. Hagberg, U. Lundqvist, and G. Persson. 1969. A proposed system of symbols for the collection of barley mutants at Svalöv. Hereditas 62:409-414.
2. Kucera, J., U. Lundqvist, and Å. Gustafsson. 1975. Inheritance of breviaristatum mutants in barley. Hereditas 80:263-278.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:476.