ITEMS FROM THE UNITED STATES

 

NEBRASKA

UNIVERSITY OF NEBRASKA AND THE USDA-ARS
Department of Agronomy, Lincoln, NE 68583, USA.


P.S. Baenziger, R.A. Graybosch (USDA-ARS), D.R. Shelton, K.S. Gill. T. Clemente, K. Arumuganathan, T. Weeks (USDA-ARS), R.C. French (USDA-ARS), D.C. Stenger (USDA-ARS), and C.J. Peterson (USDA-ARS).


1998 crop conditions.

The 1998 Nebraska wheat crop was estimated at 2,302,000 MT (84,600,000 bu), which represented a 3,160 kg/ha (47 bu/acre) state average yield on 730,000 harvest hectares (1,800,000 harvested acres). Winter wheat was planted on 770,000 hectares (1,900,000 acres). This crop had a record yield per hectare and was larger than the 1997 crop (70,300,000 bu harvested from 1,900,000 acres with a 37.0 bu/acre yield average) and the 1996 crop (73,100,000 bu harvested from 2,150,000 acres with a 34 bu/acre yield average), but smaller than the 1995 crop of 86,100,000 bu (41 bu/acre). With these high yields, the protein content was lower than normal (generally 11.5 to 12.0 %). Arapahoe continued to be the most popular and widely grown variety (28.3 % of the state) in 1998. To put Arapahoe's acceptance in perspective, it was grown on more acres in Nebraska than varieties developed by all other states and commercial seed companies combined. Alliance was the second most widely grown variety followed by Centura and Niobrara.


New germ plasm.

Culver
is a HRWW developed coöperatively by the Nebraska Agricultural Experiment Station and the USDAARS. Culver was selected from the cross 'NE82419 / Arapahoe', which was made in 1987. The pedigree of NE82419 is 'Trapper // CMN / OT /3/ CIMMYT / Scout /4/ Buckskin sib / Homestead'. Culver is an F3-derived line that was selected in the F4 generation. Culver was released primarily for its superior adaptation to and good disease resistance in dryland agriculture production systems in southcentral and southwestern Nebraska. Culver is moderately resistant to stem rust (contains Sr6 and Sr24, and other unnamed resistance genes) and leaf rust (similar to Arapahoe and most likely contains Lr16) and susceptible to WSBMV, Hessian fly, BYDV, and WSMV. Culver has a moderately low grain-volume weight, better than that of Alliance, but less than those of Arapahoe, Pronghorn, and Rawhide. The overall end-use quality characteristics for Culver should be acceptable to the milling and baking industries. Culver is recommended for growing in southern and westcentral Nebraska. In these areas, Culver is a good replacement for Alliance, because it has a longer coleoptile, better leaf rust resistance, grain-protein content, and grain-volume weight. The cultivar is genetically complementary to 2137, Alliance, Jagger, Niobrara, Pronghorn, and Windstar. Culver is not complementary to Arapahoe, Niobrara, and Vista. Like Arapahoe, Culver also seems to have a growth pattern where early spring drought can lower its performance.

Wesley is a HRWW developed by the USDA-ARS (Dr. C. J. Peterson's leadership) and the Nebraska Agricultural Experiment Station. The cultivar will be coreleased with South Dakota Agricultural Experiment Station. Wesley is a high-yielding, semidwarf wheat with adaptation characteristics similar to 2137 and is particularly well suited for production in southeast Nebraska and under irrigation. Wesley has superior straw strength, a short coleoptile, and a heading date similar to that of Scout 66. The winter hardiness of Wesley is superior to that of Scout 66 and TAM-107. Wesley is postulated to carry Sr6 and Sr17 and also has resistance to WSBMV, WSSMV, and is tolerant of acid soils. The name was chosen to honor the late Dr. John Wesley Schmidt, former wheat breeder at the University of Nebraska.

Four experimental lines (Nuplains, NE93496, NE93613, and NE 94479) are under large-scale increase for possible release in 1999. In addition, NE93405 has been held over for a third year of testing before possible release in 1999. NE93405 and NE93496 (NE85707/Thunderbird) are sister lines and are Thunderbird derivatives with along coleoptile, good winter hardiness, good test weight, large kernels, and very strong straw strength (NE93496 having the better standability). NE93613 is a sister line to Culver with a medium-length coleoptile, good winter hardiness, average test weight, medium kernels, and moderately strong straw strength. NE94479 was derived from the cross 'Arapahoe / Abilene // NE86488'. The pedigree of NE86488 is 'Colt /3/ WRR 5* / Agent // Kavkaz'. NE94479 has good winter hardiness, average test weight, medium kernels, and moderately strong straw strength. As opposed to Arapahoe and Culver, it seems to perform well despite spring droughts. Nuplains is a HWWW developed by the USDAARS (Dr. C. J. Peterson's leadership) and the Nebraska Agricultural Experiment Station. The line is derived from the cross 'Abilene / KS831872', where the pedigree of KS831872 is 'Plainsman V // Newton / Arthur 71', a high protein selection. Nuplains is a dual-purpose wheat, suitable for both bread and noodle making.


Wheat transformation and tissue culture studies.

A. Mitra, T. Clemente, S. Sato, P. S. Baenziger, R. French, and T. Weeks.

Numerous transgenic plants were created using the particle bombardment technique, and progress was made developing an Agrobacterium-facilitated method. One of the main disease-resistance efforts currently being studied is the 2-5A system, a key research area of Dr. Mitra, which may provide broad-spectrum resistance to RNA virus (such as WSMV). Mr. Todd Campbell, a graduate student, is genetically characterizing 2-5A transgenic plants. One of his more interesting findings is that that although co-transformation is common (in this research, genes on three plasmids were inserted), co-insertion is less common (i.e., the three plasmid did not always insert at the same site). The importance of this finding is that it may be possible to have genes of interest (there are two main genes in the 2-5A system) co-insert, but have selectable marker genes (often antibiotic resistance genes) insert at different genome locations that can later be bred out of the populations. Some consumers are concerned with antibiotic resistance genes in food.

The wheat transformation team at the Plant Transformation Core Research Facility at the University of NebraskaLincoln employs both microprojectile bombardment and Agrobacterium-mediated transformation protocols for the genetic engineering of wheat. Transformation efficiencies range from 26 % and 0.10.7 % on an explant/plant basis for the microprojectile and Agrobacterium-mediated protocols, respectively. Our future plans include further genetic characterization of transgenic plants, developing optimized transgenes for expression in wheat, and adding possible fungicidal genes as part of the scab initiative research.


Wheat quality research.

R.A. Graybosch, D.R. Shelton, and Gang Guo.

Functional properties of waxy wheat samples were analyzed. Wheat a -amylase activities, kernel weights and hardness values, flour Rapid Visco Analyser parameters, and falling number values were determined. These results were compared with those of the control wheats with nonwaxy starch that were planted at the same location. The comparisons showed Rapid Visco Analyser parameters, such as peak time (temperature), holding strength (trough), breakdown, setback, and final viscosity, varied significantly between the waxy and control samples (p-value = 0.0001). Waxy wheat had much lower pasting (gelatinizing) temperature, lower peak time (temperature), lower holding strength, lower setback, and final viscosity than the controls but had much higher breakdown. The falling number values of waxy wheat were much lower (about 70 seconds) than those of the controls (about 500 seconds). However, the a -amylase activities indicated that the variations in pasting properties were mainly due to the fact that the waxy starch activities of waxy wheats and control samples were approximately equal.

The T1AL·1RS wheat-rye chromosomal translocation originally found in Amigo wheat has resistance genes for stem rust, powdery mildew, and greenbug biotypes B and C, but also has a negative effect on wheat processing quality. Recently, a second T1AL·1RS translocation with Gb6, a gene conferring resistance to greenbug biotypes B, C, E, G, and I, was identified in the wheat germ plasm line GRS1201. A secalin of Mr 70 kD was found in the Amigo T1AL·1RS but did not occur in the GRS1201 T1AL·1RS. Polymorphisms detected by PCR primers derivedfrom a family of moderately repetitive rye DNA sequences also differentiated the two translocations. No differences in grain yield or grain and flour quality characteristics were observed when lines carrying 1RS from Amigo were compared to lines with 1RS from GRS1201. Hence, differences in secalin composition did not result in differential quality effects. When compared to sister lines with T1AL·1RS derived from the wheat cultivar Redland, lines with the GRS1201 had equal grain yield, but produced flours with significantly shorter mix times, weaker doughs, and lower SDS sedimentation volumes.


Personnel.

Dr. C. J. Peterson resigned from the USDA-ARS to accept a position as wheat breeder at Oregon State University. Dr. R. Graybosch will assume the role of coördinator for the USDA-ARS Hard Winter Wheat Regional Program. Mr. Delton Dovel retired after 30 years of dedicated service to the wheat improvement efforts. Mr. Pat Tenopir accepted a position with the Foundation Seed Division. Mr. Mitchell Montgomery joined the wheat improvement project in May, and Mr. Chris Hoagland joined the wheat improvement project in August as their successors. Dr. Mohammed Maroof Shah successfully completely his Ph.D. and accepted a postdoctoral position at Iowa State University.


Publications.

Baenziger PS, Moreno-Sevilla B, Peterson CJ, Shelton DR, Baltensperger DD, Haley SD, Nelson LA, McVey DV, Watkins JE, Hatchett JH, and Schmidt JW. 1998. Registration of 'Windstar' Wheat. Crop Sci 38: 894-895.

Delwiche SR, Graybosch RA, and Peterson CJ. 1999. Identification of wheat lines possessing the 1AL·1RS or 1BL·1RS wheat-rye translocation by near-infrared reflectance spectroscopy. Cereal Chem (In press).

Graybosch RA, Peterson CJ, Hareland GA, Shelton DR, Olewnik MC, He H, and Stearns MM. 1999. Relationships between small-scale wheat quality assays and commercial test bakes. Cereal Chem 76:(In press).
Graybosch RA, Lee J-H, Peterson CJ, Porter DR, and Chung OK. 1999. Genetic, agronomic and quality comparisons of two 1AL·1RS. wheat-rye chromosomal translocations. Plant Breed 118:(In press).

Graybosch RA, Peterson CJ, and Chung OK. 1999. Quality effects of rye (Secale cereale L.) Chromosome arm 1RL transferred to wheat (Triticum aestivum L.). J Cereal Sci (In press).

Wesenberg DM, Baenziger PS, Rasmusson DC, Burrup DE, and Jones BL. 1998. Registration of 88Ab536-B barley germplasm. Crop Sci 38:559.

Watkins JE, Rutledge SS, Baenziger PS, and Youngquist W. 1998. Physiologic specialization of Puccinia recondita f. sp. tritici in Nebraska during 1995 and 1996. Plant Dis 82:679-682.