ITEMS FROM THE UNITED STATES

GEORGIA / FLORIDA

UNIVERSITY OF GEORGIA
Department of Agronomy, Griffin, GA 30212, USA.

J.W. Johnson, R.D. Barnett, B.M. Cunfer, G.D. Buntin, and D.C. Bridges.

The 1998 Georgia winter wheat crop was grown on about 300,000 harvested acres, which was the lowest in 20 years. The crop production resulted in a state average yield of 44 bu/acre. Acreage was down due to wet conditions at planting. Overall, the season was characterized by a mild, wet winter followed by a dry, hot spring. Some cultivars had vernalization problems due to the warm autumn and winter and delays in planting. Cool and dry conditions prevailed through the grain-filling stage. Septoria glume blotch and leaf rust also were major factors in some low yields. Hessian fly infestations reached damaging levels in some parts of the state. Biotype L expanded its range in the state. Cereal leaf beetle also reached damaging levels this year, resulting in the spraying of several thousands acres.

Breeding.
The SRWW line GA 89482 will be increased for release. GA 89482 was selected from the cross Pioneer 2555/PF 84301//FL302, which carries the T1BL·1RS translocation. This cultivar possesses excellent resistance to powdery mildew, moderate resistance to leaf rust, and good resistance to Hessian fly. GA 89482 has excellent test weight and straw strength and is medium in maturity.

Research. Adult-plant resistance to powdery mildew was investigated for three resistant wheat cultivars, Knox 62, Massey, and Houser, and the susceptible Chancellor. Massey was the most resistant followed by Knox 62 and Houser. The resistance genes in Knox 62 are mainly additive, and seven chromosomes are involved in the adult- plant resistance. Chromosomes 5A, 7A, 4B, and 5D have genes for positive effects on resistance, whereas chromosomes 3A, 1B, and 1D have factors increasing susceptibility. The segregation of the disomic F2 populations and noncritical crosses fit a four-gene model, suggesting that chromosomes 5A, 7A, 4B, and 5D each contain one gene or one cluster of tightly linked genes.

The influence of temperature on the resistance to powdery mildew was evaluated in six wheat cultivars/lines, Axminster/8* Chancellor (Cc) (Pm1), Chul/8*Cc (Pm3b), Yuma/8*Cc (Pm4a), VPM1 (Pm4b), Kavkaz (Pm8), and Chancellor, at two temperatures of 25·C and 15·C with two isolates of powdery mildew. Resistance, based on the components of colony appearance period, infection type, colony number of unit leaf area, infected leaf area, and AUDPC, was much less effective at 15·C than at 25·C for VPM1 and 'Yuma/8*Cc', but no resistance shift was observed for Kavkaz and 'Axminster/8*Cc'. This indicated a difference of temperature-sensitivity among Pm genes or among the interactions of individual host genes with corresponding powdery mildew pathogen genes.

In recent years, Georgia wheat farmers have increasingly experienced difficulties controlling Italian (annual) ryegrass in wheat with Hoelon (diclofop). An option is to use Sencor (metribuzin) as a postemergence treatment for control of annual ryegrass, wild radish, and other broadleaf species. Some wheat cultivars are sensitive to Sencor (metribuzin), and applicationscan cause severe injury and/or stand loss. The Sencor label contains a list of known tolerant and sensitive cultivars of wheat and barley, but does not include all varieties that are currently grown in Georgia. Field trials conducted at Griffin, GA, included approximately 40 wheat cultivars that were submitted to the Statewide Variety Testing Program for testing and consideration for recommendation. Metribuzin was applied postemergence at a rate of 0.375 lb ai/acre in water at 20 GPA.

Severe wheat injury and stand reduction were observed with many cultivars, including previously known sensitive cultivars and some previously untested cultivars. Of the 37 cultivars tested, 17 were categorized as tolerant, which indicates that total crop injury did not exceed 10 %, and detectable stand reduction occurred compared to the untreated paired plot. Four cultivars were categorized as moderately tolerant, which indicates that crop injury ranged from 11 to 20 % and that stand loss was insignificant. Sixteen cultivars were categorized as sensitive, which indicates that crop injury exceeded 20 % and/or significant stand loss was observed. Seventeen of the 37 cultivars tested are currently listed as either tolerant, moderately tolerant, or sensitive on the Sencor label.

Plant Pathology. Bunt in annual ryegrass (Lolium multiflorum) seed is caused by Tilletia walkeri, a newly described Tilletia species, and was found in wheat fields at 13 locations in south central Tennessee, northern Alabama, and Georgia. Teliospores of the fungus were found at several additional sites. These collections areidentical to isolates of the smut fungus found on ryegrass seed in Oregon. Ryegrass bunt is a cryptic disease found at very low levels throughout the southeastern U.S. In all but one collection, only a few fully or partially bunted seeds were found. At one site in Georgia, about 50 % of the ryegrass seed was partially bunted, and a low percentage of fully bunted seeds was found. Even at this site, bunted ryegrass was not recognized by visual inspection of the dry seed. Ryegrass bunt was found in the same areas where smut spores were detected on wheat seed in USDA surveys in 1996-98. Ryegrass bunt is the source of spores found on wheat seed in USDA surveys in 1996 that were initially misidentified as Karnal bunt. No wheat with Karnal bunt or spores of the fungus has been found in the southeastern U.S.

Inoculum of Stagonospora nodorum was present in wheat fieldthroughout the year except in midsummer s as indicated by infection of wheat seedlings. Seedlings 17 m from the edge of a field were infected infrequently. The results suggest that inoculum travels only a short distance from wheat plants or stubble. Therefore, conidia are probably the major sources of inoculum, and it is unlikely that ascospores play a significant role in Georgia.

Entomology. The management and use of selected insecticide treatments to control aphids and suppress BYD luteovirus were studied during four seasons in winter wheat in Georgia. Aphids were mostly the bird cherry-oat aphid and greenbug during vegetative growth stages and the English grain aphid, Sitobion avenae, during grain filling. Early plantings had much greater aphid numbers and incidence of BYDV symptoms than later plantings. Planting at the end of the recommended period reduced aphid numbers and BYDV incidence and usually avoided yield losses. Treatment of seed with imidachloprid (Gaucho) at 0.5 to 1.0 oz ai/100 lb of seed reduced aphid numbers by ~ 90 % for 45 to 60 days after planting. Yield responses to Gaucho use varied with aphid number and BYDV incidence but averaged 56 bu/acre over all trials. A foliar application of lambda cyhalothrin (Karate 1E/Warrior T) on seedling or tillering wheat also controlled aphids, suppressed BYDV incidence, and prevented yield losses. Foliar applications of dimethoate reduced aphid numbers but did not reliably reduce BYDV incidence or prevent yield losses. Yield responses were highly correlated with BYDV incidence (r = 0.96) and aphid numbers (r = 0.58) during the first 60 days after planting. Economic thresholds are three aphids/30-cm row in the first 30 days after planting and five aphids/30-cm row at 30 to 60 days after planting. These results show that use of insecticides to control aphid vectors can effectively suppress BYDV incidence and minimize grain yield losses in winter wheat.

The cereal leaf beetle, Oulema melanopus, was first discovered in the late 1980s and now occurs throughout the northern two-thirds of Georgia and Alabama. Larvae can cause extensive defoliation during grain heading in winter wheat, and populations have increased to levels requiring control in the piedmont and mountain regions. Cereal leaf beetle cannot be reliably controlled by manipulating planting date or variety maturity, and plant resistance is not available in southern wheat varieties. Studies in both states during the last 4 years have examined the impact of larvae infestations on wheat yield and have examined the efficacy of controlling infestations using foliarly applied insecticides when populations exceed an economic threshold. Averaged over all trials, yield reductions have ranged from 0 to 8+ bu/acre. Most labeled insecticides are effective at killing larvae at low rates. However, M-trak, which is a Bacillus thuringiensis product, did not effectively control larvae. Nevertheless, only lambda cyhalothrin (Karate 1E/Warrior T) consistently provided season-long control with a single application when applied during egg laying before final egg hatch. Lambda cyhalothrin also controls aphids, and because it can be applied during egg laying, often can be tank mixed with foliar fungicides for leaf rust suppression. Treatment is justified when populations exceed 0.5 eggs and/or larvae per stem. Several parasites also are being released in both states to provide long-term biological control of cereal leaf beetle. The egg parasite Anaphes flavipes has been released at a number of sites in each state, and field nurseries have been established in each state to rear and release the larval parasite Tetrastichus julis. Cereal leaf beetle will continue to spread southward and increase in importance as a pest of winter wheat in Georgia and Alabama.


Publications.

Cunfer BM. 1998. Seasonal availability of inoculum of Stagonospora nodorum in the field in the southeastern U.S. Cereal Res Commun 26:259-263.

Cunfer BM and Castlebury LA. 1998. Smut fungi associated with ryegrass in wheat fields in the southeastern U.S.and their relation to Karnal bunt. Phytopathology 88:S1920.

Ge Y-F, Johnson JW, Roberts JJ, and Rajaram S. 1998. Temperature and resistance gene interactions in expression of resistance to Blumeria graminis f.sp. tritici. Euphytica 99:103-109.
Johnson JW, Ge Y-F, Cunfer BM, and Barnett RD. 1998. Adult plant resistance to powdery mildew in wheat. In: Proc 9th Inter Wheat Genet Symp (Slinkard AE ed). University Extersion Press, Saskatoon, Sasketchewan, Canada. 3:279-281.

Johnson JW, Buntin GD, Cunfer BM, Roberts JJ, and Bland DE. 1998. Registration of `GA-Luttrell' barley. Crop Sci 38:1715.

Riessen Van HW, Mask PL, Murphy JF, Cunfer BM, Buntin GD, and Johnson JW. 1998. Field resistance to the barley yellow dwarf virus-aphid complex among wheat varieties in Alabama. Phytopathology 88:S91

Shi, A.N., S. Leath, J.W. Johnson, and J.P. Murphy. 1998. Identification of powdery mildew resistance in Ae. tauschii-derived lines. In: Proc 9th Inter Wheat Genet Symp (Slinkard AE ed). University Extersion Press, Saskatoon, Sasketchewan, Canada. 3:317-319.