INSTITUTE OF AGRICULTURAL RESEARCH `SERBIA'
Center for Small Grains Kragujevac, S. Kovacevica 31, Kragujevac
34000, Yugoslavia.
Yield and quality of Yugoslav winter wheat cultivars.
Miroslav Kuburovic, Veselinka Zecevic, Desimir Knezevic, and Milanko Pavlovic.
The grain yield and some technological quality parameters
in 20 Yugoslav wheat cultivars were investigated. The experiment
was in a field at the Center for Small Grains, Kragujevac, over
a 2-year period. The results show that grain yield differed
between investigation years. The wheat cultivars Studenica, Nevesinjka,
and Kraljevica had high and stable grain yields. The investigated
cultivars had technologically good flour and dough quality. Among
the 20 cultivars, 18 belonged to the first quality class and 14
belonged to the A quality group. Cultivar KG-56S showed
the highest quality and belonged to the first quality class and
A1 quality group.
Introduction. For intensive
wheat production under diverse environmental conditions (climatic
and soil conditions) and management, the yield potential of cultivars
differs between years and localities (Djokic 1988). The yield
level depends on the genetic yield potential of a cultivar and
its yield stability in different environmental conditions (Borojevic
1981). The genetic wheat yield potential expressed in Yugoslavian
field production average from 30-40
% and in some regions almost 50 % (Borojevic et al. 1987). Thus,
there are considerable reserves for increasing the yield of winter
wheat.
Many new wheat cultivars with high yield and quality
are created in the breeding process. Technological quality is
primarily conditioned by the quality level of the cultivar, but
the realized quality is conditioned also by the agroecological
conditions of culture. The technological quality of a wheat cultivar
depends on environmental conditions and growing technology and
so requires investigating cultivars in different agroecological
conditions.
Material and methods.
Plants were grown in the field for a 2-year period at the
Center for Small Grains, Kragujevac. Twenty Yugoslav winter wheat
cultivars were investigated (see Table 1). The grain yield, sedimentation
value, moist-gluten content, quality number, and quality group
were analyzed (Table 1). The sedimentation value was determined
by the Zeleny method, and the moist-gluten content determined
by the standard hand-rinse method. Water absorption, quality
number, and quality group were determined by the Brabender farinograph
test.
Results and discussion.
Grain yield is very complex and depends on many other traits.
Environmental conditions have a significant influence on grain
yield (Borojevic 1981). Grain yield differs according to cultivar
characteristics, technology level of culture, and environmental
conditions. A breeding program may take many years to create
a cultivar that is high-yielding and adapted to certain conditions
and field management (Mak Key 1966).
Many of the cultivars investigated had higher grain yields in 1995 than in 1996. Kraljevica, Francuska, Kompas, and Evropa had the highest grain yield in 1995. The highest-yielding cultivars in 1996 were Rodna, Studenica, and Jugoslavija. Those cultivars had a significantly higher grain yield than the experimental average at a 1 % probability. Cultivars NS Rana-5, Nevesinjka, and Slavija had significantly higher grain yields (at a 5 % probability) than the average of the experiment in the first or second years. Studenica, Nevesinjka, and Kraljevica had high, stable yields, which were significantly higher than the average of experiments in both experimental years. There were significant differences in grain yield among cultivars in each year, which agrees with Kuburovic et al. (1988). The cultivars Jugoslavija, Francuska, and Kompas yielded higher than average for these 2 years, and these results agree with those of Mladenov (1996).
Table 1. Grain yield of 20 Yugoslavian winter wheat cultivars grown in a field at the Center for Small Grains, Kragujevac, over a 2-year period (1995-96).
Cultivar | Grain yield (T/ha) | ||
---|---|---|---|
1995 | 1996 | Average | |
Jugoslavija | 6.84 | 6.94** | 6.89 |
Francuska | 7.25 | 6.56 | 6.90 |
Kompas | 7.18** | 6.36 | 6.77 |
Beogradjanka | 6.10 | 5.46 | 5.78 |
Zemunka | 6.24 | 6.76 | 6.50 |
Studenica | 6.94 | 7.00** | 6.97 |
Levcanka | 5.70 | 6.78 | 6.24 |
KG-56 | 6.54 | 6.22 | 6.38 |
Srbijanka | 6.14 | 6.26 | 6.20 |
Evropa | 7.19** | 6.36 | 6.78 |
NS Rana 5 | 7.06** | 6.04 | 6.55 |
Nevesinjka | 7.10* | 6.85* | 6.97* |
Slavija | 7.20* | 6.51 | 6.85 |
Danica | 6.78 | 6.16 | 6.46 |
Proteinka | 7.14** | 5.97 | 6.56 |
Rodna | 6.22 | 7.04** | 6.63 |
Kraljevica | 7.34** | 6.64 | 7.00* |
PKB-20 | 6.16 | 6.15 | 6.15 |
Takovcanka | 6.96 | 6.76 | 6.86 |
Dicna | 6.39 | 6.32 | 6.35 |
Average | 6.72 | 6.46 | 6.59 |
Cultivar (A) | Year (B) | Interaction(AB) | |
LSD 5% | 0.332 | 0.468 | 0.469 |
1% | 0.438 | 0.618 | 0.619 |
* Significant at P = 0.005.
** Significant at P = 0.001.
The quality parameters investigated show that all
the cultivars had good technological-quality flour and dough.
The sedimentation value indicate that all of the investigated
cultivars belonged to the first quality class, except for Kompas
and Zemunka, which belonged to the second quality class. Eighteen
of the investigated cultivars had a moist-gluten content above
35 %. Nevesinjka (27.73 %) and Rodna (29.56 %) had the lowest
values for this trait. The results were similar for water absorption.
All of the investigated cultivars had water absorption above
60 %, except Nevesinjka (59.0 %) and Rodna (59.0 %). The cultivar
KG-56S was the highest in quality and belonged to the first
quality class and the A1 quality group. The cultivar
PKB-20 also belonged to A1 quality group but had
lower values than KG-56S for the other quality parameters
(Table 2). These results agree with our previous investigations
(Pavlovic et al. 1994).
Table 2. Sedimentation, moist gluten, water absorption, quality number and quality group for 20 Yugoslav winter wheat cultivars grown in 1995-96 at the Center for Small Grains, Kragujevac.
Cultivar | Sedimentation value (ml) | Moist-gluten content (%) | Water absorption (%) | Quality number | Quality group |
---|---|---|---|---|---|
Jugoslavija | 57 | 39.45 | 65.2 | 71.3 | A2 |
Francuska | 55 | 37.18 | 60.0 | 78.0 | A2 |
Kompas | 36 | 45.55 | 61.8 | 51.2 | B2 |
Beogradjanka | 45 | 40.54 | 63.4 | 80.4 | A2 |
Zemunka | 30 | 37.98 | 61.8 | 44.5 | C1 |
Studenica | 55 | 36.80 | 63.9 | 71.6 | A2 |
Levcanka | 58 | 37.76 | 64.4 | 76.0 | A2 |
KG-56S | 69 | 43.50 | 64.8 | 86.4 | A1 |
Srbijanka | 52 | 38.97 | 63.0 | 66.7 | B1 |
Evropa | 53 | 42.36 | 61.8 | 74.6 | A2 |
NS Rana 5 | 65 | 41.12 | 62.4 | 72.2 | A2 |
Nevesinjka | 62 | 27.73 | 59.0 | 65.4 | B1 |
Slavija | 62 | 35.73 | 63.7 | 80.0 | A2 |
Danica | 59 | 39.63 | 63.4 | 83.2 | A2 |
Proteinka | 55 | 38.22 | 67.4 | 65.2 | B1 |
Rodna | 67 | 29.56 | 59.0 | 71.6 | A2 |
Kraljevica | 45 | 40.15 | 64.4 | 60.6 | B1 |
PKB-20 | 55 | 38.05 | 61.8 | - | A1 |
Takovcanka | 62 | 39.25 | 62.2 | 71.9 | A2 |
Dicna | 62 | 34.67 | 62.2 | 80.4 | A2 |
Conclusion. The results
show that grain yield is independent of the year of the investigation.
The cultivars Studenica, Nevesinjka, and Kraljevica had high
and stable grain yields. The investigated cultivars had good
technological-quality flour and dough. Of the 20 investigated
cultivars, 18 belonged to the first quality class, and 14 belonged
to the A quality group. The cultivar KG-56 S had the highest
quality and belonged to the first quality class and A1
quality group.
References.
Borojevic S. 1981. Principles and methods of plant
breeding. Novi Sad.
Borojevic S and Misic T. 1987. Cultivar as a factor
of improvement of wheat production. Wheat 6 million tons. Novi
Sad. p. 15-28.
Djokic A.. 1988. The some important traits and
yield of Zemun's
wheat cultivars. The improvement of wheat production and other
small grains in the aim of creating of strategy of technological
development of Yugoslavia. Kragujevac. p. 39-60.
Kuburovic M, Stojanovic S, and Rafailovic M. 1988.
The analysis of some quantitative traits of wheat (Triticum
aestivum ssp. vulgare L) in different environmental
conditions. Rev Work Inst for Small Grains, Kragujevac 9:15-26.
Mak Key J. 1966. The wheat plant as a model in
adaptation to high productivity under different environments.
Contemporary Agriculture pp. 11-12.
Mladenov N. 1996. The study of genetic variability
of wheat line and cultivars in different environmental conditions.
Ph.D. thesis. University of Belgrade, Agricultural faculty.
Pavlovic M, Kuburovic M, Zecevic V, and Knezevic
D. 1994. Parameters of technological quality and components
of yield of some winter wheat varieties. J Cereal Flour Technol
21(4):94-96.
UNIVERSITY OF LLEIDA AND INSTITUTE FOR FOOD AND AGRICULTURAL RESEARCH AND TECHNOLOGY (UdL-IRTA)
Center of Research & Development, Rovira Roure 177, 25198-Lleida,
Spain.
National production.
The total area of wheat grown in Spain has varied
very little in the last 3 years. However, the area devoted to
durum wheat continued to increase, because of E.U. subsidies for
each cultivated hectare (see Table 1).
Table 1. Total area and yield for the Spanish bread and durum wheat crops for the years 1993-96.
Crop Season | Bread wheat | Durum wheat | ||
---|---|---|---|---|
Area (x 103 ha) | Yield (x 103 T) | Area (x 103 ha) | Yield (x 103 T) | |
1993-94 | 1,370.4 | 3,381.4 | 599.3 | 920.0 |
1994-95 | 1,459.3 | 2,565.6 | 633.8 | 392.1 |
1995-96 | 1,374.7 | 4,468.1(est.) | 650.7 | 1,705.3* |
* estimate.
C. Royo.
Dual-purpose triticale. A research project studied the suitability of dual-purpose triticale (forage and grain in the same cropping season). The effect of clipping the crop at the end of tillering on grain yield, grain quality and plant growth was studied in winter and spring triticales.
Grain shrivelling. The
effect of drought and terminal water-stress simulated by
a senescing agent (potassium iodide, KI) on triticale yield, yield
components, and grain filling was studied in Trujillo triticale
and four derived NILs.
Morphophysiological traits in durum wheat.
Work is presently underway to identify morphophysiological traits
related to drought resistance in durum wheat.
J. Lloveras, A. Lopez-Querol, and J.A. Martin-Sanchez.
The cultivar IRTA 2009 was included in the
Spanish official list in 1996. This variety (pedigree: CM 85295
(PFAU "S"/Seri//Bow"S")
was selected from CIMMYT material (entry No. 109 of 8th SAWSN).
Compared to Anza, IRTA 2009 is slightly taller and later heading,
but with similar yield. However, IRTA 2009 has the high quality
required by the Spanish standards (W about 300 and P/L 1.5-2.0).
Durum wheat and triticale program.
C. Royo.
A triticale breeding program operated from 1981 to
1996. The following five spring varieties were released: Trujillo
(1987), Piky (1991), Tupi
(1993), Tabo
(1993), and Terrola (1994). Both farmers and industry recently
have become less interested in triticale. Thus, fewer resources
have been allocated to triticale research and more to other species,
i.e., durum wheat. The observed decrease in the area devoted
to triticale in Spain is mainly because of the expansion of the
area planted to durum wheat (a subsidized crop) in some areas
of the country. Durum wheat hectarage in 1996 was three times
that of 1990. Consequently, our breeding program is now focussed
on durum wheat, in to response to the increasing demand for new
varieties adapted to Spanish climactic conditions. The durum
varieties Valira (1990), Bolo (1993), Borli
(1994), and Bolido (1996) have been released. Our breeding methodology
is based on individual plant selection. The main goals are grain
quality, adaptability, stability, and yield potential.
Quality.
Work on the quality of durum wheat is targeted in
two main areas: studying the of effect of the T1B-1R
translocation on the yield and quality in durum wheat (J. Zarco
and A. Michelena), and looking at the effect of nitrogen fertilization
in grain quality of durum and bread wheat (A. Abad, J. Lloveras,
and A. Michelena).
Personnel.
Dr. A. Michelena left his duties for durum wheat
breeding to become responsible for the of Department of Quality.
Dra. C. Royo assumed the responsibility of durum wheat and triticale
breeding. Ing. J. Zarco, formerly of CIMMYT research group, is
beginning Ph.D research.
Publications.
Araus JL, Santiveri P, Bosch-Serra D, Royo C, and Romagosa I. 1992. Carbon isotope discrimination in ear parts of triticale: influence on grain filling. Plant Physiol 100:1033-1035.
Garcia
del Moral LF, Boujenna A, Insa JA, Arbonaes
A, and Royo C. 1994. Evaluation of a set of triticale genotypes
for dual-purpose (forage and grain). Melhoramento 33(1):291-296.
Michelena A, Romagosa I, Martin-Sanchez
JA, and Lopez-Querol
A. 1995. Influencia ambiental y varietal en diferentes parametros
de calidad y rendimiento en trigo duro. Investigacion
Agraria. Produccion
y Proteccion
Vegetales. Vol 10 (2)191-201.
Michelena A, Romagosa I, Martin-Sanchez
JA, and Lopez-Querol
A. 1995. Estabilidad de la calidad y el rendimiento del trigo
harinero. Inv Agraria Prod y Prot Veg Vol 10 (2) 203-213.
Ramos JM, Garcia
del Moral LF, Boujenna A, Serra J, Insa JA, and Royo C. 1996.
Grain yield, biomass and leaf area of triticale in response to
sowing date and cutting stage in three Mediterranean contrasting
environments. J Agric Sci Cambridge. 126:253-358.
Royo C. 1992. El triticale: bases para el cultivo
y aprovechamiento. Ed Mundi Prensa, Madrid. 96 pp.
Royo C. 1997. Yield components as affected by forage
removal in winter and spring triticale. Grass and Forage Sci
(In press).
Royo C, Abaza M, Cantero C, Caldero
A, Ramos JM, and Garcia
del Moral LF. 1996. Likening between the effect of drought and
terminal water-stress simulated by a senescing agent in triticale.
J Agron Crop Sci 176:31-38.
Royo C, Insa JA, Boujenna A, Ramos JM, Montesinos
E, and Garcia
del Moral LF. 1994. Yield and quality of spring triticale for
forage and grain as influenced by sowing date and cutting stage.
Field Crops Res 37(3):161-168.
Royo C, Montesinos E, Molina-Cano JL, and Serra
J. 1993. Triticale and other small grain cereals for forage
and grain in Mediterranean conditions. Grass and Forage Sci
48:11-17.
Royo C and Paraes
D. 1996. Yield and quality of winter and spring triticales for
forage and grain. Grass and Forage Sci 51:449-455.
Royo C, Penella E, Tribo
F, and Molina-Cano JL. 1996. Aptitude of spring and winter
triticales for dual purpose (forage and grain) in Mediterranean
conditions. In: Triticale: Today and Tomorrow. (Guedes-Pinto
H, Darvey N, Carnide VP eds). Kluwer Academic Publishers, The
Netherlands. pp. 843-849.
Royo C, Rodriguez A, and Romagosa I. 1993. Differential
adaptation of complete and substituted triticale. Plant Breed
111:113-119.
Royo C and Romagosa I. 1996. Effect of a forage
removal at the first node detectablestage on the growth of winter
and spring triticale. Grass and Forage Sci 51:170-179.
Royo C, Romagosa I, and Rodriguez A. 1991. Comparative
adaptation of triticale and spring wheat in Spain. In:
Proc Second Inter Triticale Symp. CIMMYT, Maexico.
pp. 593-598.
Royo C, Soler C, and Romagosa I. 1995. Agronomical
and morphological differentiation among winter and spring triticales.
Plant Breed 114:413-416.
Royo C and Tribo
F. 1997. Triticale and barley for grain and for dual-purpose
(forage + grain) in a Mediterranean-type environment. I.
Growth analyses. Aust J Agric Res 48:in press.
Royo C and Tribo
F. 1997. Triticale and barley for grain and for dual-purpose
(forage + grain) in a Mediterranean-type environment. II.
Yield, yield components and quality. Aust J Agric Res 48:in
press.
UNIVERSIDAD POLITICNICA DE MADRID.
Departamento Biotecnologia,
E.T.S.Ingenieros Agronomos,
Ciudad Universitaria, 28040 -
Madrid, Spain.
A. Delibes and I. Lopez-Brana.
CONSEJO SUPERIOR DE INVESTIGACIONES CIENTIFICAS.
Serrano 115. 28006 -
Madrid, Spain.
D. Romero and M.F. Andres.
UNIVERSIDAD DE LLEIDA AND INSTITUT DE RECERCA I TECNOLOGIA AGROALIMENTARIES (UdL-IRTA).
Rovira Roure, 177. 25198 -
Lleida, Spain.
J.A. Martin-Sanchez, E. Sin, C. Martinez, and A. Michelena.
Transfer of genes for resistance to Heterodera avenae, from
two wild grasses: use in wheat breeding programs.
Two sources of resistance to H. avenae have
been extensively used (see AWN Vol. 41, 1995), the Cre2
gene from Ae. ventricosa and an as yet unnamed gene from
Ae. triuncialis. Available data on segregation for resistance
data are compatible with a simple monogenic dominant inheritance.
Both sources are effective against the different H. avenae
pathotypes (Table 1).
Table 1. Sources of resistance to Heterodera avenae and their resistance to the different known pathotypes (R = resistant, MR = moderately resistant, and S = susceptible).
Source of resistance | Pathotype | ||||||
---|---|---|---|---|---|---|---|
Ha11 Fr3 | Ha41 Fr1 | Ha12 Fr2-4 | Ha71 (Spain) | Torralba (Spain) | N. Harene Hg1 | Etelhem Hg3 | |
Ae. triuncialis | R | ? | R | R | R | R | R |
Cr1 (Loros/Aus 10894) | R | R | R | R | R | MR-S | MR-S |
Cr2(Ae. ventricosa) | R | R | R | R | R | S | S |
UNIVERSIDAD POLITICNICA DE MADRID.
Departamento Biotecnologia,
E.T.S.Ingenieros Agronomos,
Ciudad Universitaria, 28040 -
Madrid, Spain.
A. Delibes and I. Lopez-Brana.
UNIVERSIDAD DE LLEIDA AND INSTITUT DE RECERCA I TECNOLOGIA AGROALIMENTARIES (UdL-IRTA).
Rovira Roure, 177. 25198 -
Lleida, Spain.
J.A. Martin-Sanchez,
E. Sin, C. Martinez,
and A. Michelena.
Servicio de Investigacion
Agraria, Finca La
Orden, 06187 - Guadajira,
Badajoz, Spain.
J. del Moral and A. Mejias.
Transfer of a gene for resistance to Hessian fly (Mayetiola
destructor) from Aegilops ventricosa to cultivars of wheat.
Hessian fly is an important pest in wide areas of
the Guadiana (Badajoz) and the Guadalquivir (Cadiz,
Cordoba,
Sevilla) valleys in southern Spain. This pest has caused severe
losses during the several years in these areas.
The GP biotype seems to be present in the area.
The genetic variability of this biotype currently is being study
using molecular markers. Subsequent comparative studies will
be made with samples of Hessian fly from other areas of the Iberian
Peninsula. A number of agronomic studies aimed at developing
cultural techniques that may interfere with pest development also
are being studied.
Concurrently, the transfer of a resistance gene from
Ae. ventricosa to wheat cultivars has began. The gene
(H27) is located in chromosome 4Mv and is linked
to an acid phosphatase isozyme marker, Acph-Mv1.
This marker can facilitate the transfer of the resistance gene
into cultivars.
Publications.
Bleve-Zacheo T, Melillo MT, Andraes
MF, Zacheo G, and Romero D. 1995. Ultrastructure of initial
response of graminaceous roots to infection by Heterodera
avenae. Nematologica 41:80-97.
Delibes A, Del Moral J, Martin-Sanchez JA, Mejias A, Gallego M, Casado D, Sin E, and Lopez-Brana I. 1996. Hessian flyresistance gene transferred from chromosme 4Mv of Aegilops ventricosa to Triticum aestivum. Theor Appl Genet (In press).
Delibes A, Romero MD, Martin-Sanchez
JA, Lopez-Brana
I, Duce A, Sin E, Andraes
MF, Martinez
C, and Aguaded S. 1995. Obtencion
de cultivares resistentes al nematodo Heterodera avenae
por transferencia gaenica
desde especies de Aegilops. Phytoma Espana
73:22-25.
del Moral J, Mejias A, Chacon A, Delibes A, Lopez-Brana I, Martin-Sanchez JA, Sin E, and Martinez C. 1994. Consideraciones para el desarrollo de un programa de sanidad respecto a la plaga Mayetiola destructor Say en los trigales de Badajoz. I Congreso de la Sociedad Espanola de Agricultura Ecologica. Toledo, September, 1994.
del Moral J, Mejias
A, Chacon
A, Delibes A, Lopez-Brana
I, Martin-Sanchez
JA, Sin E, and Martinez
C. 1995. Resistencia a Mayetiola destructor Say
de una coleccion
de variedades y lineas
de trigo. V Jornadas Cientificas
de la Sociedad Espanola
de Entomologia
Aplicada. Sevilla, 20-24
November, 1995.
del Moral J, Mejias
A, Chacon
A, Delibes A, Lopez-Brana
I, Martin-Sanchez
JA, Sin E, and Martinez
C. 1995. Localizacion
cromosomica
de la resistencia a Mayetiola destructor Say en
una linea
de trigo.
Lopez-Brana
I. 1996. The random amplified polymorphic DNA (RAP) technique
as a tool to study the variability of Heterodera avenae.
Cereal Cyst Nematode Workshop. In: III Inter Nemat Congr.
Guadalupe 7-9
July, 1996.
Lopez-Brana
I, Romero D, and Delibes A. 1996. Analysis of Heterodera
avenae populations by the random amplified polymorphic
(RAPD) technique. Genome 39:118-122.
Martin-Sanchez
JA, Delibes A, Andraes
MF, Duce A, Lopez-Brana
I, Martinez
C, Michelena A, Romero D, and Sin E. 1996. New Source of resistance
to cereal cyst nematode (Heterodera avenae Woll).
5th Inter Wheat Conf. 10-14
June, 1996, Ankara, Turkey. p. 143.
Melillo MT, Bleve-Zacheo T, Delibes A, Lopez-Brana
I, Romero D, and Andraes
MF. 1995. Histochemical localization of enzymes related to Heterodera
avenae resistance in wheat. Nematologica 41:321.
Romero MD, Andraes
MF, Balague JM, Delibes A, Duce A, Lara A, Lopez-Brana
I, Martin-Sanchez
JA, Martinez
C, Michelena A, and Sin E. 1996. Las especies de Aegilops
como fuentes de resistencia al nematodo de los cereales Heterodera
avenae.-Resumenes. VII Congreso Nacional de la Sociedad
Espanola
de Fitopatologia.
p 67.
Romero MD, Andraes
MF, Balague JM, Delibes A, Duce A, Lara A, Lopez-Brana
I, Martin-Sanchez
JA, Martinez
C, Michelena A, and Sin E. 1996. Aegilops triuncialis,
a source of resistance to Heterodera avenae (CCN).
III International Nematology Congress, 7-12
July. Guadalupe. Abstract.
Romero MD, Andraes
MF, Lopez-Brana
I, and Delibes A. 1996. A pathogenic and biochemical comparison
of two Spanish populations of cereal cyst nematode. Nematol Medit
24:235-244.
Romero D, Delibes A, Lopez-Brana
I, Mena M, Duce A, and Andraes
MF. 1995. Studies of the chromosomic location of a gene conferring
resistance to Heterodera avenae transferred from
the wild grass Aegilops ventricosa to hexaploid
wheat. Nematologica 41:337.
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