TRIGEN SEED SERVICES LLC.

8024 Telegraph Rd., Bloomington, MN 55478, USA.

Robert W. Romig.

Despite a few growing pains and untimely rain that affected some planting operations, we successfully transferred winter wheat segregating populations between the Northern and Southern Hemispheres this year. We had a relatively successful nursery in Argentina that was planted in early June with seed harvested in the U.S. in early May. Selections harvested there in late November were planted in south Texas in mid-December, where they are growing well.

We had good germination in the straight winter populations harvested in Argentina. Presumably, a couple of cold-shock cycles prior to planting, each lasting 2 days at below freezing temperatures, were adequate to break any germination dormancy that may have been present. We have our fingers crossed, hoping that these will be vernalized to produce seed this spring. The real proof of the usefulness of this `shuttling concept' will be if we can successfully repeat the cycle this year.

Our crossing program in Chile turned out well again. However, we experienced a delay in receiving the seed because of a delivery mix-up in Miami. Consequently, the F1s were planted a bit later than expected. Germinated seeds from the `winter x winter' crosses are being chilled under refrigeration to develop vernalized F1 seedlings for transplanting later this spring.

This year's program currently consists of a nursery in Texas that has segregating lines and winter and spring wheat trials and a nursery near Yuma, AZ, with new F1s, segregating lines derived from last year's nursery in Minnesota along with a number of wheat and triticale increase plots. Work is in progress to establish spring wheat trials this year near Moses Lake, WA.

We have changed the organizational basis of the company this year (1995) from a sole proprietor enterprise to a limited liability company. This is a relatively new type of organization that offers certain useful financial protection features for small business organizations such as ours. However, we continue to follow the original operating and breeding strategies, reported in last year's Newsletter.

Our business strategy is evolving. The central concepts are to be a provider of improved wheat genetics, to bring these to market by working through local or regional producers/marketers, and to form synergistic alliances with other breeding programs.


ITEMS FROM ARGENTINA

Institute of Biological Resources

Castelar, Argentina.

Chromosome mapping: comparison between recombination frequency values of centromeric genes obtained by using telocentric and normal chromosomes.

G. Tranquilli, L. Pfluger, S. Lewis, and E. Suarez.

The telocentric lines developed in Chinese Spring by Sears have been very useful tools in chromosome mapping, mainly in gene location on specific chromosome arms, as well as their linkage with the centromere. Nevertheless, it was proposed (Sears 1972) that the chromosome pairing in the centromere proximal area and, consequently, crossing over, are reduced in a heteromorphic bivalent. Therefore, the obtained recombination frequency in these cases is lower than that expected when normal bivalents are present. In order to analyse this situation, the recombination frequencies of two centromeric genes, one on each chromosome arm of 6B, are being evaluated using both telocentric or normal chromosomes.

The considered marker genes are Amp-B1 (aminopeptidase) on 6BS and B2 (awn inhibitor) on 6BL. The Argentinean cultivar Sinvalocho MA carries a silent allele for Amp-B1 and the b2 allele (awned). On the other hand, Chinese Spring codifies for Amp-B1 and possesses the B2 allele (awnless). Chinese Spring (euploid) and its double ditelocentric line for chromosome 6B were crossed with the 6B substitution line of Sinvalocho MA (Sin) into Chinese Spring (CS(Sin6B)). Both F1s (2n = 42 and 2n = 41 + t6BS + t6BL, respectively) were backcrossed with CS(Sin6B), and the resultant offspring classified according to the following: karyotype, aminopeptidase pattern, and presence or absence of awns. A total of 140 and 188 individuals corresponding to each BC1, respectively, have been evaluated. In the "telocentric chromosome system", individuals with abnormal karyotypes were taken into account if they could be classified unambiguously as parental or recombinant type. Up to the present, no recombinant has been found considering Amp-B1 and the centromere. On the other hand, for the B2 locus, a further generation will be obtained in order to classify unambiguously the gene expression, because many plants, from both BC1s, showed an intermediate phenotype for awn expression.

Reference.

Sears E. 1972. Reduced proximal crossing-over in telocentric chromosomes of wheat. Genet Iber 24:233-239.

Genetic variability of an Argentinean land race of wheat

E. Suarez, M.L. Appendino, G. Tranquilli, and L. Bullrich.

Through the evaluation of a wheat land race grown by a farmer in the province of Catamarca (Argentina), nine lines were isolated showing variation for the following features: grain colour (from white to dark); glumes (pubescent and glabrous); awnedness (awned, semiawned, and awnless); Erysiphe graminis tritici reaction (susceptible and resistant); ear emergence time (an extreme difference of 15 days, for plants cultivated in long days); coleoptile (purple and green); and plant height. This last character (plant height) showed a range of variation of about 50 cm, a difference similar to that observed between semidwarf and tall varieties carrying Norin genes. Nevertheless, all lines were responsive to gibberellic acid. The results for these eight traits showed a high potential of genetic variability, even for characters not necessary present in commercial cultivars.

On the other hand, when the nine lines were isoenzymatically assayed, no variability was found for IBF (iodine binding factor), a very polymorphic system among commercial cultivars.


Cordoba National University

College of Agriculture, P.O. Box 509, Cordoba, 5000, Argentina.

Biological yield, harvest index, and ear weight as indirect selection criteria for grain yield in wheat.

W. Londero, V. Pantano, F. Bidinost, D. Bonelli, R. Roldon, and R. Maich.

The objective of this study was to determine the effect of indirect selection for grain yield using biological yield (BY), harvest index (HI), and ear weight (PSP) as selection criterion. During 1990, 160 crosses (single and double) in the first selfing generation were cultivated at Ferreyra (Cordoba). A random sample of 800 S1 plants was taken from them and characterized through BY, HI, and PSP. For each selection criteria, a group of 15 S1 superior plants was created. At the same time, the best 15 were selected according to the phenotypic value (Ph). The 60 S1 derived lines were evaluated for grain yield with two sowing dates (May and June) during 3 years (1991/92/93). Analysis of variance of the experiment was conducted according to the factorial model. Nonsignificant differences were obtained among criteria, but a significant (P < 0.05) `selection criteria x sowing date' interaction existed. For the first sowing date, the material selected for Ph yielded more than the HI selections. For the second, the material selected for BY performed worse than those selected for Ph and HI.

Correlated response to recurrent selection for grain yield in wheat.

F. Salvagiotti, N. Conton, C. Ferraris, B. Ferro, C. Olmos, and R. Maich.

The objective of this work was to measure the efficiency of a recurrent selection program for grain yield and to determine the effect on associated characters. During 1993, 54 S1:2 lines, 17 from each selection cycle (C0, C1, and C2), were evaluated at Ferreyra (Cordoba) for two sowing dates (May and June) and without replication per date. Grain yield (GY), biological yield (BY), and harvest index (HI) were recorded for each plot. Nonsignificant differences were obtained among cycles for GY and BY, but not for HI. For HI, the C1S1:2 and C2S1:2 lines showed higher values than C0S1:2. The C1 and C2 group mean differences were not significant.

Prospects of wheat production in accordance with an integrated point of view: farmer, technician, and contractor.

R. Maich, P. Daher, and M. Bianchi.

A networking point of view and farmer, technician, and contractor interactions are responsible for the success of agriculture in marginal areas. To make the reconversion effective, the farmer must choose suitable cultivars and farm practices, the technician must put more emphasis on his plant breeding programs, and the contractor must provide capital for technological development.

Influence of early-generation selection for proteins on yield and bread making quality in wheat.

G. Manera, Z. Gaido, and M.E. Dubois.

The lack of wheat that meets millers' specifications in the central semiarid region of Argentina indicates the necessity to promote the development of materials adapted to great fluctuations in rainfall. These materials should have high potential production, stability in yields, and good commercial and industrial characteristics. The objective of this work was to study the effect of early generation selection for high protein content on yield and bread making quality in wheat grains available for the semiarid region. Yield and bread making quality measured as proteins (P), humid gluten (GH), dry gluten (GS), and mixogram (M) was determined in 45 lines of wheat. These lines, selected in the F3 generation for high protein content (selection pressure = 15 %), came from three selection environments (semiarid, intermediate, and humid), reproduced in the semiarid. No significant correlation occurred between protein content and yield in any of the three studied environments; all resulted in good bread making quality (xP = 14.42; xGH = 35.08; xGS = 13.13; and xM = 6.33). These results indicate the convenience of simultaneous selection in early generations, for yield and protein content in wheat grown for semiarid regions.

Variation of the epidermal characters of the flag leaf in experimental and commercial bread wheat (Triticum aestivum L.) varieties sown late in the season.

Ana L. Pascualides, S. Patricia Gil, Maroa M. Cerana, and Laura Gonzolez.

The goal of this study is to determine the anatomical adaptations of the flag leaf blade in Triticum aestivum L. varieties to different, local, environmental conditions. The commercial varieties, PROINTA Oasis and Las Rosas INTA, and the experimental ones, T7 and T24, were sown in plots of 11 m x 45 m at two different spacings in the row (15 cm and 30 cm) late in the season. A random design with factorial treatment was used for this test. Temporary slices of the abaxial and adaxial leaf epidermis were made in order to characterize them and to calculate the stomatal frequency (SF), the stomatal index (SI), the frequency of silico-suberose pairs, and the frequency of prickles (PF). A further analysis shows that the distance factor does not influence the variables for the abaxial face. But, in the adaxial face, both SF and PF are influenced by distance, whereas the SI does not change. The number of stomata/mm2 increases at 15 cm spacing for every cultivar, except PROINTA Oasis, which has a different behavior. The same trend occurs with PF.