From MASWheat:
Grain protein content (GPC) is one of the most important quality factors for pasta and bread wheats. Breeding for an increase in GPC is difficult because the genetic variation for this character is small compared to variations caused by the environment, besides there is a negative correlation between GPC and grain yield. The stripe rust resistance gene Yr36 is closely linked to the Gpc locus, and the same markers used for tagging Gpc-B1 can be used for Yr36.
Genes and alleles updated with links to MASWheat and the Wheat Gene Catalog at Komugi:
Gpc-B1 (Triticum) , Gpc-B1a (Triticum), and Gpc-B1b (Triticum)
Note that curated genes and alleles in GrainGenes contain the genus in parenthesis within the name.
QTL updated with links to the WGC. Note that grain protein content qtl have been published as QPro and QGpc and are in the 'protein content' and 'grain protein content' trait categories. All are available on CMap.
Prasad et al. (2003) QTL analysis for grain protein content using SSR markers and validation studies using NILs in bread wheat. TAG-106-659
QGpc.ccsu-2B.1
QGpc.ccsu-2D.1
QGpc.ccsu-3D.1
QGpc.ccsu-3D.2
QGpc.ccsu-7A.1
Khlestkina et al. (2009) A new gene controlling the flowering response to photoperiod in wheat. EUP-165-579
QGpc.ipk.7B
Gonzalez-Hernandez et al. (2004) Mapping genes for grain protein concentration and grain yield on chromosome 5B of Triticum turgidum (L.) var. dicoccoides. EUP-139-217
QGpc.ndsu-5B.1
QGpc.ndsu-5B.2
QGpc.ndsu-5B.3
Joppa et al. (1997) Mapping gene(s) for grain protein in tetraploid wheat (Triticum turgidum L.) using a population of recombinant inbred chromosome lines. CRS-37-1586
QGpc.ndsu-6B
QGpc.ndsu-6Ba
QGpc.ndsu-6Bb
Groos et al. (2003) Genetic analysis of grain protein-content, grain yield and thousand-kernel weight in bread wheat. TAG-106-1032
QPro.inra-2A
QPro.inra-3A
QPro.inra-4D
QPro.inra-7D
Blanco et al. (1996) Quantitative trait loci influencing grain protein content in tetraploid wheats. PBR-115-310
QPro.mgb-4B
QPro.mgb-5A
QPro.mgb-6A.1
QPro.mgb-6A.2
QPro.mgb-6B
QPro.mgb-7B
Blanco et al. (2002) Detection of grain protein content QTLs across environments in tetraploid wheats PMB-48-625
QPro.mgb-7A